A new colosteid, Deltaherpeton hiemstrae gen. et sp. nov., is described from the Mississippian Upper Viséan site at Delta, Iowa. Deltaherpeton is represented by a skull roof and both jaws. The new taxon is unique among colosteids in having an internasal and single midline postparietal. An additional midline pair of cf. ‘interfrontonasals’ may be present. Characters previously used to define the colosteids are reviewed and a refined diagnosis for the family Colosteidae is presented. Synapomorphies which unite Deltaherpeton, Colosteus, Greererpeton, and Pholidogaster as Colosteidae are: premaxilla with fang pair; dentary with notch for receipt of premaxillary fang; mandible with single elongate exomeckelian fenestra; pre-narial infraorbital lateral line terminating at ventral margin of premaxilla just anterior to external naris; and post-narial infraorbital lateral line terminating at the ventral margin of the maxilla just posterior to the external naris. Our review of dermal bones in the colosteid snout concludes that no specimen is sufficiently free of distortions or breakage to indicate clearly whether or not the prefrontal borders the external naris, or that an anterior tectal is present. The morphology of Deltaherpeton and the revised data presented for colosteids do not clarify the relationship of colosteids to other early tetrapods.

Among hyoliths from the Givetian Kersadiou Formation in northwestern France one form is assigned to Hyolithes triliratus Hall, 1879, which is used as the basis for Brittanella new genus. The occurrence of this species in France supports affinity for fauna in that region to the Eastern Americas Realm. Other hyoliths from this unit include one species of Hyolithes Eichwald, which is left in open nomenclature, and one conch and one platyclaviculate operculum that cannot be identified to genus. The occurrence of these fossils in concretions in an oxygen-stressed dark gray shale supports the notion that this environment served as a refuge for taxa that were less efficient competitors with other normal marine benthos in the Devonian.

Fossilized leaf mines and other traces of phytophagous insects provide a unique window into ecological and evolutionary associations of the past. Leaf-mining flies (Diptera: Agromyzidae) are an important component of the recent leaf-mining fauna, but their fossil record is sparse compared to other mining insect lineages; many putative agromyzid body fossils and traces are dubiously assigned. Agromyzid leaf mines often can be distinguished from those of other insects by the presence of an intermittent, fluidized frass trail that may alternate between the sides of the mine. Here, we describe two new Paleogene leaf mine fossils, Phytomyzites biliapchaensis Winkler, Labandeira and Wilf n. sp. from the early Paleocene of southeastern Montana, USA, occurring in leaves of Platanus raynoldsii (Platanaceae); and Phytomyzites schaarschmidti Wappler n. sp., from the middle Eocene of Messel, Germany, occurring in leaves of Toddalia ovata (Rutaceae). These fossils both exhibit frass trails indicative of an agromyzid origin, and P. biliapchaensis also exhibits associated stereotypical marks identical to damage caused by feeding punctures of extant adult female Agromyzidae prior to oviposition. Phytomyzites biliapchaensis represents the earliest confirmed record of Agromyzidae, and one of the earliest records for the large dipteran clade Schizophora. Plant hosts of both species belong to genera that are no longer hosts of leaf-mining Agromyzidae, suggesting a complex and dynamic history of early host-plant associations and, for the early Paleocene example, an evolutionary, possibly opportunistic colonization in the midst of the ecological chaos following the end-Cretaceous event in North America.

Re-evaluation of teeth referred to the Late Cretaceous shark Ptychodus polygyrus from North America are hereby determined to be synonymous with another species having similar tooth morphology, Ptychodus marginalis Agassiz. Ptychodus marginalis differs from P. polygyrus by having an elevated and rounded tooth crown with transverse ridges restricted centrally on the apex of the crown with a clearly defined marginal area. The type specimen of P. marginalis was described from Middle Cenomanian to Middle Turonian deposits in the English Chalk, and teeth possessing identical tooth morphologies are found within contemporary units in North America, whereas teeth having the typical P. polygyrus morphology are restricted to Late Santonian-Early Campanian deposits.

Triassic lungfish (Dipnoi) have been extensively documented from the Gondwanan continental and marine shelf deposits of Africa and Madagascar (Teixeira, 1949; Lehman et al., 1959; Beltan, 1968; Martin, 1979, 1981; Kemp 1996), Australia (Kemp, 1993, 1994, 1997a, 1998), India (Jain et al, 1964; Jain, 1968), and Antarctica (Dziewa, 1980). Numerous records also exist from Laurasian landmasses including Europe (Agassiz, 1838; Schultze, 1981), North America (Case, 1921) and central and eastern Asia (Liu and Yeh, 1957; Vorobyeva, 1967; Martin and Ingavat, 1982). By comparison, nothing is known of contemporary lungfish fossils from the Middle East. Thus, the recent recovery of a single tooth plate representing a new geographic occurrence of the genus Ceratodus Agassiz, 1838 from paralic marine deposits of the Jilh Formation, a latest Anisian to lower Carnian unit that crops out along the eastern margin of the Proterozoic Arabian Shield in central Saudi Arabia (Fig. 1), is significant because it provides the stratigraphically oldest record of dipnoans from the Arabian Peninsula.

A large (maximum length 80 mm), tubular, corset-like problematic fossil from the early Cambrian (Cambrian Series 2, Stage 3) Sirius Passet Lagerstätte of North Greenland is interpreted as the lorica of an ancestral loriciferan. in addition to the double circlet of 7 plates composing the lorica, Sirilorica carlsbergi new genus, new species also preserves up to six multicuspidate cuticular denticles that are similar in shape to the pharyngeal teeth of priapulid worms, although their location is suggestive of scalids. Whilst traditionally placed as a sister group of priapulid worms within Vinctiplicata (Scalidophora), recent molecular sequence data suggest that loriciferans might be more closely related to nematomorphs. the limited morphological information available from Sirilorica is consistent with this interpretation, placing the Sirius Passet fossil within the total-group of Loricifera, within the Loricifera + Nematomorpha clade.

A specimen of the aquatic reptile Champsosaurus sp. from the Paleocene Black Peaks Formation in southwestern Texas is the southernmost yet known. the fragmentary specimen exhibits some unusual features, such as a great anterior extent of the quadratojugal on the lower temporal arch, and cannot be attributed with confidence to any of the named species. Champsosaurus appears to have been tolerant of temperate climates and had a northern latitudinal range exceeding that of crocodylians. It seems likely that the brief southward extension in range of Champsosaurus during early Paleocene time resulted from a decrease in mean annual temperature, comparable to over 10° of paleolatitude.

An exceptionally well-preserved silicified bivalve from the Upper Permian of Texas is described and assigned to a new genus and species, Cassiavellia galtarae, placed in the family Bakevelliidae. The species represents one of the earliest and best characterized unequivocal occurrences of the multivincular ligament in the superfamily Pterioidea. The silicified material provides a wealth of information on the morphology of inadequately known Paleozoic pterioideans, including hitherto undescribed aspects of the larval shell, auricular sulcus, muscle scars, and dental ontogeny. The discovery of the condyle-fossa complex on the anteroventral shell margin, a feature previously undescibed in Bivalvia, raises the question of the homology and taxonomic significance of the problematic subumbonal ridge-like structures in Pterioidea. In life, C. galtarae was probably an epifaunal right-pleurothetic bivalve, byssally attached to hard or raised flexible substrata. In addition to C. galtarae, another new species, C. nadkevnae, is placed in Cassiavellia.

A fusuline fauna consisting of 9 species of 4 genera from the Xiala Formation of the Mujiucuo section, Xainza County, Tibet, China is described. The fusuline fauna is dominated by Nankinella and Chusenella and indicates a Midian (Late Guadalupian) age. The earliest record of fusuline fauna during the Midian in the Lhasa Block suggests that the block rifted later than the Qiangtang Block to the north and the Baoshan and Tengchong blocks to the east, all of which yield much earlier fusuline faunas of Yakhtashian (Artinskian) age, but had drifted away from Gondwana to a relatively warm temperate zone in the Late Guadalupian (Middle Permian).

A rare new turban-shaped basal Middle Cambrian gogiid eocrinoid, Turbanicystis inflatus new genus and species is described from the Kaili Formation (Kaili Biota), Guizhou Province, China. It co-occurs with three other markedly different gogiids, Sinoeocrinus, Globoeocrinus, and Balangicystis, and is part of a unique Middle Cambrian community that is dominated by echinoderms.

Two abdominal molts representing a small cymothoid isopod are described from the marine late Miocene Gram Formation of southern Denmark. Abdominal molts are generally regarded as being difficult or even impossible to assign to genus or family due to overlap of morphological characters. A preliminary study supported by statistical and multivariate analysis of the genera most closely resembling the specimens reveals that although some overlap in characters do occur, the genera can to a large extent be defined by a diagnostic set of features. The Danish Miocene taxon described herein is assigned to Aega Leach, 1815 as the only known fossil of this important cosmopolitan genus.

Representatives of the family Strephocladidae have been considered as fossil relatives (i.e., stem-group) of Mantodea (mantises) based on characters of the forewing morphology. Here we describe new specimens from the Wellington Formation that we assign to the strephocladid species Homocladus grandis Carpenter, 1966. The range of morphological variation exhibited by the new material, in addition to wing morphology variability documented in extant mantises and roaches, suggest that H. ornatus Carpenter, 1966 and Paracladus retardatus Carpenter, 1966, reported from the same formation, are new junior subjective synonyms of H. grandis. We describe the first hind wing for this species based on a well-preserved specimen. It exhibits a combination of character states unique to dictyopteran insects.

An enigmatic microproblematicum, Tuxekanella Riding and Soja (1993), has been identified in the shallow water carbonate succession of the Late Silurian Skala formation (Podolia, Ukraine). the newly found specimens suggest most probably the algal nature of this enigmatic microfossil. Tuxekanella is known only from two extremely distant regions at the moment: the Alexander terrane in Alaska and Podolia. the possibility of their paleobiogeographical link is discussed in the context of paleotectonic models, assuming the peri-Gondwanan origin of the Alexander Terrane.

The rudist bivalve Durania aguilae Adkins was described from the Mendez Formation in Mexico and originally attributed to the Santonian. We revise this species and show that it belongs to the genus Chiapasella Müllerried. We also recognize that Chiapasella aguilae (Adkins) is a senior synonym of Chiapasella trechmanni Mitchell and Gunter, an important biostratigraphic indicator in the late late Maastrichtian of Jamaica. Chiapasella aguilae is therefore an important late late Maastrichtian marker fossil that has been recorded from Jamaica, Mexico, and Cuba.

Clark and Martin (1901), in their study of the Paleogene deposits of Maryland, figured drawings of a fragment of a spatangoid echinoid from the Lower Eocene Nanjemoy Formation and a broken echinoid spine from the Upper Paleocene Aquia Formation. Echinoids are an important element of Paleogene faunas in the eastern United States, but these are the only reported occurrences of this age from either Maryland or Virginia and are therefore significant to the understanding of the evolution and paleogeography of echinoids in this region. The figured specimens were attributed to collections at Johns Hopkins University, but without catalog numbers. During study of the paleobiology collections of the National Museum of Natural History (NMNH), Smithsonian Institution, two vials were discovered with small, handwritten labels referencing the figure numbers in Clark and Martin (1901). The contained specimens were recognized as those figured in the report. They have been cataloged in the NMNH collections as follows:

A rudist fauna composed of the radiolitids Potosites tristantorresi new genus, new species and Radiolites acutocostata (Adkins), the hippuritids Barrettia cf. ruseae Chubb, Torreites sanchezi (Douvillé), and Vaccinites vermunti Mac Gillavry, plagioptychids, and antillocaprinids is described from a rudist limestone succession in the central Mexican State of San Luis Potosí. The previously known species, one radiolitid from Texas and three hippuritids from the Caribbean Biogeographic Province, indicate the early Campanian. The new genus is a large radiolitid with a coiled and canaliculated left valve. New observations on R. acutocostata and V. vermunti shell morphology and variability are provided. Observations made on well preserved specimens of the latter species clarify the outer shell layer structure of the hippuritids right valve and has taxonomic implications. This is the first report of genus Torreites in Mexico and one of the northernmost records of genus Barrettia, already reported in southern Mexico from Chiapas State. This rudist limestone succession (informally ‘Temazcal limestones’ herein) is an easy-to-recognize cartographical unit; up to now it and Santonian and Turonian units have been mapped as the El Abra Formation in most available geological maps. Recognition of the successive carbonate platform intervals, between El Abra and Cardenas formations, improves the geological mapping and the understanding of the Mexican Gulf western margin evolution during the Late Cretaceous.

An assemblage of seventeen species of Small Shelly Fossils, dominated by the brachiopod Eothele tubulus and species of the mollusk Yochelcionella, is described from the basal Kinzers Formation of Thomasville, Pennsylvania. The occurrence extends southwards the distribution of an Early Cambrian fauna (Cambrian Series 2, Stage 4) that is otherwise characteristic of the eastern shelf of Laurentia from New York to Greenland. The poorly known acrothelid brachiopod Eothele tubulus is redescribed based on large collections of ventral valves. The shell structure of E. tubulus is characterized by orthogonal baculae, and represents the oldest known example of a baculate shell structure, indicating that this type of shell structure evolved already in the Early Cambrian.

Recent field work at the Deseadan (late Oligocene) locality of Quebrada Fiera (Mendoza Province, Argentina) has provided new fossil remains that greatly increased the faunal assemblage of this site. A good number of specimens correspond to the Family Archaeohyracidae (Notoungulata) and are presented in this paper. Most of them are recognized as a unique taxon, corresponding to the genus Archaeohyrax, characterized by the presence of a strong, shallow, wide sulcus on the labial side of the talonid of m3. At specific level, it is identified with the species A. suniensis previously recognized at Salla (Bolivia). Specimens with different tooth wear degree show an intraspecific variation in occlusal morphology comparable to that observed among the Bolivian material. In addition to A. suniensis, an isolated p2 is tentatively related to Archaeotypotherium, due to their hypsodont but rooted condition and larger size. Two other isolated lower molars from this site had been previously related to this genus but they are comparable to the new specimens ascribed to Archaeohyrax. The identification of Archaeotypotherium is not well supported at present, but if later confirmed, it would increase the temporal distribution of the genus, so far recorded in Tinguirirican (early Oligocene) localities of Chile and Argentina. The recognition of A. suniensis in Quebrada Fiera establishes a greater affinity with the lower latitude Deseadan fauna of Salla than with that from Patagonia, where the genus is represented by A. patagonicus. Other faunal elements from Quebrada Fiera, however, do not support this affinity with Bolivia. Xenarthrans or some marsupials are equivalent to those from Patagonia, while other taxa such as the recently described notohippid or a rare, tiny metatheria are at present exclusive for Mendoza. The increasing knowledge of the Quebrada Fiera assemblage contributes to a better understanding of the paleogeographic scenario of Southern South America during late Oligocene.

An Anchitherium specimen with a nearly complete series of the upper cheek teeth (P2–M3), from the upper part (ca.17–18 Ma) of the Hiramaki Formation, Kani Basin, Gifu Prefecture, Japan, was previously referred to Anchitherium “hypohippoides” nomen dubium. Despite that it provides one of the best examples of significant dental characters of Asian Anchitherium, it has remained undescribed and unprepared until recently. Although a paucity of materials from Asia makes the taxonomy of Asian Anchitherium difficult to assign, comparison showed that the specimen should be reassigned to Anchitherium aff. A. gobiense; it differs from A. aurelianense and is rather similar to A. gobiense from China by virtue of large size and expanded hypostyles. The Japanese Anchitherium also shows distinct features including straight (flattened) ectolophs with narrow mesostyles, rudimentary crochets, and enamel protuberances at the lingual mouth of the median valleys. This combination of accessory features has not been known in Asian Anchitherium and seems to be rarely observed among the diversified European species. The existence of Japanese Anchitherium implies early species diversification in East Asia that predates a greater diversification in Europe associated with the Middle Miocene Climatic Optimum and supports paleogeographical and paleozoological connection to the Asian mainland under a warm and humid climate prior to the formation of the Japanese archipelago (ca. 16.5 Ma).

New field collections allow the study and description of Valvata juliae new species from the Pliocene upper Burgi Member of the Koobi Fora Formation of Kenya. The shell morphology of this species varies from trochospiral to planispiral to open coiled. The species is restricted to a short stratigraphic interval. Valvata juliae is considered as an invader of the Turkana Basin during a lacustrine transgression event. The open coiling of the species is interpreted as an ecophenotypic response to a high level of environmental stress caused by lake level fluctuations and emergence of delta systems. These environmental conditions brought Valvata juliae to extinction soon after it invaded the Turkana Basin.