Hostname: page-component-76fb5796d-5g6vh Total loading time: 0 Render date: 2024-04-26T08:41:47.873Z Has data issue: false hasContentIssue false

South American occurrences of the Adamanian (Late Triassic: Latest Carnian) index taxon Stagonolepis (Archosauria: Aetosauria) and their biochronological significance

Published online by Cambridge University Press:  14 July 2015

Andrew B. Heckert
Affiliation:
New Mexico Museum of Natural History and Science, 1801 Mountain Road Northwest, Albuquerque 87104,
Spencer G. Lucas
Affiliation:
New Mexico Museum of Natural History and Science, 1801 Mountain Road Northwest, Albuquerque 87104,

Abstract

Aetosaur fossils from the Upper Triassic Ischigualasto Formation of Argentina formerly assigned to Aetosauroides scagliai Casamiquela and Argentinosuchus bonapartei Casamiquela are reassigned to Stagonolepis robertsoni Agassiz (small specimens) and S. wellesi (Long and Ballew) (larger specimens). Numerous features of the skull, vertebral column, appendicular skeleton, and particularly the armor, of these Argentinian aetosaurs are identical to those of Stagonolepis and differ from other aetosaurs. Identification of Stagonolepis in South America has important implications for the correlation of nonmarine strata across Late Triassic Pangea. Stagonolepis is an index taxon of the Adamanian land-vertebrate faunachron of latest Carnian age. Its occurrences in the Ischigualasto and Santa Maria formations, as well as at the type locality in the Lossiemouth Sandstone of Scotland and in the Blasensandstein of the German Keuper, cross-correlate with a well-established tetrapod biostratigraphy of the Chinle Group in western North America. Thus, all Stagonolepis records are of Adamanian (latest Carnian) age, not Otischalkian (early or “middle” Carnian) age, as proposed by some earlier workers. This correlation also demonstrates that the oldest known dinosaurs are not from the Ischigualasto and Santa Maria formations. Previous workers obtained radioisotopic dates of 227.8 Ma from the Ischigualasto Formation, providing a maximum numerical date for Stagonolepis-bearing units. This date must be late Carnian, not of Ladinian age, and thus supports the apparent age of the Ladinian-Carnian boundary (232 Ma) of time scales based on the Newark Supergroup in eastern North America.

Type
Research Article
Copyright
Copyright © The Paleontological Society

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Agassiz, L. 1844. Monographie des Poissons Fossiles du Vieux Grés Rouge ou Systéme Dévonien (Old Red Sandstone) des Iles Britanniques ed de Russie. Jent et Gassman, Neuchâtel, 171 p.Google Scholar
Alcober, O., and Parrish, J. M. 1997. A new poposaurid from the Upper Triassic of Argentina. Journal of Vertebrate Paleontology, 17:548556.CrossRefGoogle Scholar
Benton, M. J. 1990. Origin and interrelationships of dinosaurs, p. 1130. In Weishampel, D. B., Dodson, P. P., and Osmólska, H. (eds.), The Dinosauria. University of California Press, Berkeley.Google Scholar
Benton, M. J., and Clark, J. M. 1988. Archosaur phylogeny and the relationships of the Crocodylia, p. 295338. In Benton, M. J. (ed.), The Phylogeny and Classification of the Tetrapods, Volume 1. Clarendon Press, Oxford.Google Scholar
Bonaparte, J. F. 1966. Chronological survey of the tetrapod-bearing Triassic of Argentina. Brevoria, 251:113.Google Scholar
Bonaparte, J. F. 1967. Dos nuevas “faunas” de reptiles Triasicos de Argentina. Gondwana Symposium Proceedings and Papers, 1:283306.Google Scholar
Bonaparte, J. F. 1971a. Los tetrapodos del sector superior de la Formacion Los Colorados, la Rioja, Argentina (Triásico Superior) I Parte. Opera Lillona, 22:1183.Google Scholar
Bonaparte, J. F. 1971b. Annotated list of the South American Triassic tetrapods. Gondwana Symposium Proceedings and Papers, 2:665682.Google Scholar
Bonaparte, J. F. 1978. El Mesozoico de America del Sur y sus tetrapods. Opera Lilloana, 26:1596.Google Scholar
Cabrera, A. 1944. Sobre un estegocéfaol del Triásico Argentino. Notas Museo de la Plata, 9:421429.Google Scholar
Casamiquela, R. M. 1960. Notica preliminar sobre dos nuevos estagonlepoideos Argentinos. Ameghiniana, 2:39.Google Scholar
Casamiquela, R. M. 1961. Dos nuevos estagonolepoideos Argentinos de Ischigualasto, San Juan. Revista Asocíacion Geológia Argentina, 16:143203.Google Scholar
Casamiquela, R. M. 1967. Materiales adicionales y reinterpretación de Aetosauroides scagliai (de Ischigualasto, San Juan). Revista del Museo de La Plata, Paleontologia Serie 5, 33:173196.Google Scholar
Case, E. C. 1932. A perfectly preserved segment of the armor of a phytosaur, with associated vertebrae. Contributions from the Museum of Paleontology, University of Michigan, 4:5780.Google Scholar
Cox, C. B. 1965. New Triassic dicynodonts from South America, their origins and relationships. Philosophical Transactions of the Royal Society of London B, 248:457516.Google Scholar
Desojo, J. 1999. Los aetosaurios (Reptilia, Archosauria) Sudamericanos: Su status taxonómico. Ameghiniana, 36 (supplement to no. 4):10R.Google Scholar
Gradstein, F. G., Agterberg, F. P., Ogg, J. G., Hardenbol, J., van Veen, P., Thierry, J., and Huang, Z. 1995. A Triassic, Jurassic and Cretaceous time scale, p. 95126. In Berggren, W. A., Kent, D. V., Aubry, M.-P., and Hardenbol, J. (eds.), Geochronology, Time Scales and Global Stratigraphic Correlation. SEPM Special Publication 54, Tulsa.CrossRefGoogle Scholar
Harland, W. B., Armstrong, R. L., Cox, A. V., Craig, L. E., Smith, A. G., and Smith, D. G. 1990. A Geologic Time Scale 1989. Cambridge University Press, Cambridge, 263 p.Google Scholar
Heckert, A. B., and Lucas, S. G. 1996. Revision of the South American Aetosaur (Archosauria: Pseudosuchia) record with implications for the absolute age of the Late Triassic Chinle Group, USA. Geological Society of America Abstracts with Programs, 28(7):365.Google Scholar
Heckert, A. B., and Lucas, S. G. 1999. A new aetosaur from the Upper Triassic of Texas and the phylogeny of aetosaurs. Journal of Vertebrate Paleontology, 19:5068.CrossRefGoogle Scholar
Heckert, A. B., and Lucas, S. G. 2000a. Taxonomy, phylogeny, biostratigraphy, biochronology, paleobiogeography, and evolution of the Late Triassic Aetosauria (Archosauria: Crurotarsi). Zentralblatt für Geologie und Paläontologie Teil I 1998 Heft 11-12, p. 15391587.Google Scholar
Heckert, A. B., and Lucas, S. G. 2000b. Global correlation of the Triassic theropod record. Gaia, 15:6374.Google Scholar
Huene, F. v. 1942. Die fossilen Reptilien de südamerikanischen Gondwanalandes, ergebnisse der Sauriergrabungen in Südbrasilien 1928/1929. C.H. Beck'Sche Verlagsbuchhandlung, München.Google Scholar
Hunt, A. P., and Lucas, S. G. 1991a. The Paleorhinus biochron and the correlation of the non-marine Upper Triassic of Pangaea. Palaeontology, 34:487501.Google Scholar
Hunt, A. P., and Lucas, S. G. 1991b. A new rhynchosaur from West Texas (USA) and the biochronology of Late Triassic rhynchosaurs. Palaeontology, 34:191198.CrossRefGoogle Scholar
Hunt, A. P., Lucas, S. G., Heckert, A. B., Sullivan, R. M., and Lockley, M. G. 1998. Late Triassic dinosaurs from the western United States. Geobios, 31:511531.CrossRefGoogle Scholar
Huxley, T. H. 1859. On the Stagonolepis robertsoni (Agassiz) of the Elgin Sandstone; and on the recently discovered footmarks in the Sandstones of Cummingstone. Proceedings of the Geological Society of London, 15:440460.CrossRefGoogle Scholar
Huxley, T. H. 1875. On Stagonolepis robertsoni, and on the evolution of the Crocodile. Proceedings of the Geological Society of London, 31:423438.CrossRefGoogle Scholar
Huxley, T. H. 1877. The crocodilian remains found in the Elgin Sandstones, with remarks on the ichnites of Cummingstone. Memoirs of the Geological Survey of the United Kingdom, Monograph 3, 58 p.Google Scholar
Kent, D. V., and Olsen, P. E. 1999. Astronomically tuned geomagnetic polarity time scale for the Late Triassic. Journal of Geophysical Research, 104:1283112841.CrossRefGoogle Scholar
Kent, D. V., and Olsen, P. E. 2000. Implications of an astronomically timed chronology for the Late Triassic. Zentralblatt für Geologie und Paläontologie Teil I 1998(7-8).Google Scholar
Kent, D. V., Olsen, P. E., and Witte, W. K. 1995. Late Triassic-earliest Jurassic geomagnetic polarity sequence and paleolatitudes from drill cores in the Newark rift basin, eastern North America. Journal of Geophysical Research, 100:1496514998.CrossRefGoogle Scholar
Langer, M. C., and Schultz, C. L. 2000. A new species of the Late Triassic rhynchosaur Hyperodapedon from the Santa Maria Formation of south Brazil. Palaeontology, 43:633652.CrossRefGoogle Scholar
Long, R. A., and Ballew, K. L. 1985. Aetosaur dermal armor from the Late Triassic of southwestern North America, with special reference to material from the Chinle Formation of Petrified Forest National Park. Museum of Northern Arizona Bulletin, 47:4568.Google Scholar
Long, R. A., and Murry, P. A. 1995. Late Triassic (Carnian and Norian) tetrapods from the southwestern United States. New Mexico Museum of Natural History and Science Bulletin 4, 254 p.Google Scholar
Lucas, S. G. 1998. Global Triassic tetrapod biostratigraphy and biochronology. Palaeogeography, Palaeoclimatology, Palaeoecology, 143:347384.CrossRefGoogle Scholar
Lucas, S. G. 2000. Tetrapod-based correlation of the nonmarine Triassic. Zentralblatt für Geologie und Paläontologie, 1998(7-8):497521.Google Scholar
Lucas, S. G., and Heckert, A. B. 1996. Late Triassic aetosaur biochronology. Albertiana, 17:5764.Google Scholar
Lucas, S. G., and Heckert, A. B. 2001. The aetosaur Stagonolepis from the Upper Triassic of Brazil and its biochronological significance. Neues Jahrbuch für Geologie und Paläontologie Monatsheft, 2001:719732.CrossRefGoogle Scholar
Lucas, S. G., and Hunt, A. P. 1993. Tetrapod biochronology of the Chinle Group (Upper Triassic), western United States. New Mexico Museum of Natural History and Science Bulletin, 3:327329.Google Scholar
Lucas, S. G., Hunt, A. P., and Long, R. A. 1992. The oldest dinosaurs. Naturwissenschaften, 79:171172.CrossRefGoogle Scholar
Lydekker, R. 1887. The fossil Vertebrata of India. Records of the Geological Survey of India, 20:5980.Google Scholar
Nicholson, J. A., and Lydekker, R. 1887. A manual of palaeontology for the use of students, with a general introduction on the principles of palaeontology. Nature, London, 1624 p.Google Scholar
Novas, F. E. 1993. New information on the systematics and postcranial skeleton of Herrerasaurus ischigualastensis (Theropoda: Herrerasauridae) from the Ischigualasto Formation (Upper Triassic) of Argentina. Journal of Vertebrate Paleontology, 13:400423.CrossRefGoogle Scholar
Novas, F. E. 1996. Dinosaur monophyly. Journal of Vertebrate Paleontology, 16:723741.CrossRefGoogle Scholar
Parrish, J. M. 1986. Locomotor adaptations in the hind limb and pelvis of the Thecodontia. Hunteria, 1(2):335.Google Scholar
Parrish, J. M. 1994. Cranial osteology of Longosuchus meadei and the phylogeny and distribution of the Aetosauria. Journal of Vertebrate Paleontology, 14:196209.CrossRefGoogle Scholar
Reig, O. A. 1959. Primeros datos descríptivos sobre nuevos reptiles arcosaurios del Triásico de Ischigualasto (San Juan, Argentina). Revista Associacion Geologica de Argentina, 13:257270.Google Scholar
Reig, O. A. 1961. Acerca de la posición sistemática de la familia Rauisuchidae y del género Saurosuchus (Reptilia, Thecodontia). Publicaciones de la Museo Cincia Naturale Mar del Plata, 1:73114.Google Scholar
Reig, O. A. 1963. La presencia de dinosaurios saurisquios en los “Estratos de Ischigualasto” (Mesotriásico Superior) de las provincias de San Juan y La Rioja (República Argentina). Ameghiniana, 3:320.Google Scholar
Rogers, R. R., Swisher, C. C. III, Sereno, P. C., Monetta, A. M., Forster, C. A., and Martínez, R. N. 1993. The Ischigualasto tetrapod assemblage (Late Triassic, Argentina) and 40Ar/39Ar dating of dinosaur origins. Science, 260:794797.CrossRefGoogle ScholarPubMed
Romer, A. S. 1960. Vertebrate-bearing continental Triassic strata in Mendoza Region, Argentina. Geological Society of America Bulletin, 71:12791294.CrossRefGoogle Scholar
Romer, A. S. 1962a. La evolucion expolosiva de los rhynchosaurios del Triasico. Revista Museo Argentino Ciencia Natural Ciencia Zoologia, 8(1):114.Google Scholar
Romer, A. S. 1962b. The fossiliferous Triassic deposits of Ischigualasto, Argentina. Breviora, 156:17.Google Scholar
Sereno, P. C. 1993. The pectoral girdle and forelimb of Herrerasaurus ischigualastensis . Journal of Vertebrate Paleontology, 13:425450.CrossRefGoogle Scholar
Sereno, P. C., and Novas, F. E. 1992. The complete skull and skeleton of an early dinosaur. Science, 258:11371140.CrossRefGoogle ScholarPubMed
Sereno, P. C., and Novas, F. E. 1993. The skull and neck of the basal theropod Herrerasaurus ischigualastensis . Journal of Vertebrate Paleontology, 13:451476.CrossRefGoogle Scholar
Sereno, P. C., Forster, C. A., Rogers, R. R., and Monetta, A. M. 1993. Primitive dinosaur skeleton from Argentina and the early evolution of Dinosauria. Nature, 361:6466.CrossRefGoogle Scholar
Walker, A. D. 1961. Triassic reptiles from the Elgin area: Stagonolepis, Dasygnathus, and their allies. Philosophical Transactions of the Royal Society of London B, 244:103204.Google Scholar
Zacarias, J. D. 1981. Una nova especie de tecodonte aetosaurio (Aetosauroides subsulcatus, sp. nov.) de Foramacáo Santa Maria, Triassico de Rio Grande du Sul, Brasil. Unpublished M.S. thesis, University Federal of the state of Rio Grande, Porta Alegre, Brasil, 69 p.Google Scholar