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Upper Steptoean (Upper Cambrian) Trilobites from the Mckay Group of Southeastern British Columbia, Canada

Published online by Cambridge University Press:  11 August 2017

Brian D. E. Chatterton
Affiliation:
Department of Earth and Atmospheric Sciences, University of Alberta, Edmonton, Alberta T6G 2E3, Canada
Rolf Ludvigsen
Affiliation:
Denman Institute of Research on Trilobites, 3390 Denman Road, Denman Island, British Columbia VOR 1T0, Canada

Abstract

An abundant and well preserved trilobite fauna is described from Upper Cambrian calcareous mudstones of the McKay Group, near Cranbrook in southeastern British Columbia. The trilobites are mostly articulated, consisting of similar numbers of molts and carcasses. They are representative of a new deep water biofacies, the Wujiajiania Biofacies, and a new Wujiajiania sutherlandi Fauna of late Steptoean age. The trilobites were collected from a narrow interval (<20 m thick) of richly fossiliferous strata, in a thick sequence of unfossiliferous to sparsely fossiliferous strata of similar lithology. The fauna includes fourteen species, six of which are new: Aciculolenus palmeri, Burnetiella leechi, Hedinaspis canadensis, Labiostria westropi, Pterocephalia norfordi, and Wujiajiania sutherlandi.

Type
Research Article
Copyright
Copyright © 1998, The Paleontological Society 

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References

Aitken, J. A. 1966. Middle Cambrian to Middle Ordovician cyclic sedimentation, southern Rocky Mountains of Alberta. Bulletin of Canadian Petroleum Geology, 14:405441.Google Scholar
Aitken, J. A. 1971. Control of Lower Paleozoic sedimentary facies by the Kicking Horse Rim, southern Rocky Mountains, Canada. Bulletin of Canadian Petroleum Geology, 19:557569.Google Scholar
Apollonov, M. K., and Chugaeva, M. N. 1983. Some trilobites from the Cambro-Ordovician boundary deposits at Batyrbay Gorge, Lesser Karatau, p. 6690. In Apollonov, M. K., Bandaletov, S. M., and Ivshin, N. K. (eds.), The Lower Paleozoic Stratigraphy and Paleontology of Kazakhstan. Kazakhstan SSR Academy of Sciences, K. I. Satpaev Institute of Geological Sciences. (In Russian.).Google Scholar
Apollonov, M. K., Chugaeva, M. N. and Dubinina, S. V. 1984. Trilobites and Conodonts of the Batyrbay Section (Uppermost Cambrian-Lower Ordovician) in Malyi Karatau Range, Atlas of the Palaeontological Plates). Nauka, Kazakh SSR Publishing House, Alma Ata, 48 p.Google Scholar
Belt, T. 1867. On some new trilobites from the Upper Cambrian rocks of North Wales. Geological Magazine, 4:294295.Google Scholar
Bright, R. C. 1959. A paleoecologic and biometric study of the Middle Cambrian trilobite Elrathia kingii (Meek). Journal of Paleontology, 33:8398.Google Scholar
Burmeister, H. 1843. Die Organisation der Trilobiten, aus ihren lebenden varwandten Entwickelt; nebst einer systematischen Übersicht aller zeither beschriebenen Arten. Reimer, Berlin, 147 p.Google Scholar
Chatterton, B. D. E., Johanson, Z., and Sutherland, G. 1994a. Form of the trilobite digestive system: alimentary structures in Pterocephalia . Journal of Paleontology 68:294305.CrossRefGoogle Scholar
Chatterton, B. D. E., Edgecombe, G. D., Speyer, S. E., Hunt, A. S., and Fortey, R. A. 1994b. Ontogeny and relationships of the Trinucleoidea (Trilobita). Journal of Paleontology, 68:523540.Google Scholar
Chien, Yi-yuan. 1961. Cambrian trilobites from Sandu and Duyun, southern Kweichow. Acta Paleontologica Sinica, 9:91129.Google Scholar
Clarkson, E. N. K., and Taylor, C. M. 1995. Ontogeny of the trilobite Olenus wahlenbergi Westergård, 1922 from the upper Cambrian Alum Shales of Andrarum, Skåne, Sweden. Transactions of the Royal Society of Edinburgh: Earth Sciences, 86:1334.Google Scholar
Evans, C. S. 1933. Brisco-Dogtooth Map-Area, British Columbia. Geological Survey of Canada Summary Report. 1932. Part A II:106176.Google Scholar
Fortey, R. A. 1974. The Ordovician trilobites of Spitsbergen I. Olenidae. Norsk Polarinstitutt Skrifter, 160, 129 p.Google Scholar
Fortey, R. A. 1975. The Ordovician trilobites of Spitsbergen II. Asaphidae, Nileidae, Raphiophoridae and Telephinidae of the Valhallfonna Formation. Norsk Polarinstitutt Skrifter, 162, 207 p.Google Scholar
Fortey, R. A. 1985. Pelagic trilobite as an example of deducing the life habits of extinct arthropods. Transactions of the Royal Society of Edinburgh, 76:219230.CrossRefGoogle Scholar
Fortey, R. A. 1990. Ontogeny, hypostome attachment and trilobite classification. Palaeontology, 33:529576.Google Scholar
Fortey, R. A., and Chatterton, B. D. E. 1988. Classification of the trilobite Suborder Asaphina. Palaeontology, 31:165222.Google Scholar
Fortey, R. A., and Owens, R. M. 1991. The Early Ordovician trilobite Beltella . Proceedings of the Bristol Naturalists' Society (1989), 49:6979.Google Scholar
Fortey, R. A., and Wilmot, N. 1991. Trilobite cuticle thickness in relation to palaeoenvironment. Paläontologische Zeitschrift, 65:141151.CrossRefGoogle Scholar
Gabrielse, H., and Yorath, C. J. 1992. Geology of the Cordilleran Orogen in Canada. Geology of Canada, Number 4, Geological Survey of Canada, 844 p.Google Scholar
Hall, J., and Whitfield, R. P. 1877. Paleontology. In King, C. Report of the Geological Exploration of the Fortieth Parallel. United States Army Engineer Department Professional Paper, 4:197302.Google Scholar
Henningsmoen, G. 1957. The trilobite family Olenidae. With description of Norwegian material and remarks on the olenid and Tremadocian series. Skrifter utgitt av Det Norske Videnskaps-Akademi i Oslo Matematisk-naturvidenskapelig klasse 1957, 303 p.Google Scholar
Henningsmoen, G. 1975. Moulting in trilobites. Fossils and Strata, 4:179200.Google Scholar
Hohensee, S. R., and Stitt, J. H. 1989. Redeposited Elvinia Zone (Upper Cambrian) trilobites from the Collier Shale, Ouachita Mountains, west-central Arkansas. Journal of Paleontology, 63:857879.CrossRefGoogle Scholar
Howell, B. F. 1935. Cambrian and Ordovician trilobites from Herault, southern France. Journal of Paleontology, 9:222238.Google Scholar
Jaekel, O. 1909. Über die Agnostiden. Zeitschrift der deutschen geologischen Gesselschaft, 61:380401.Google Scholar
Jell, P. A. 1978. Trilobite respiration and genal caeca. Alcheringa, 2: 251260.CrossRefGoogle Scholar
Jell, P. A. 1989. Some aberrant exoskeletons from fossil and living arthropods. Memoirs of the Queensland Museum, 27:491498.Google Scholar
Jell, P. A., Hughes, N. C., and Brown, A. V. 1991. Late Cambrian (post Idmaean) trilobites from the Higgins Creek area, western Tasmania. Memoirs of the Queensland Museum, 30:455485.Google Scholar
Kobayashi, T. 1935. The Briscoia fauna of the late Upper Cambrian in Alaska, with descriptions of a few Upper Cambrian trilobites from Montana and Nevada. Japanese Journal of Geography and Geology, 12:3957.Google Scholar
Kobayashi, T. 1938. Upper Cambrian fossils from British Columbia with a discussion on the isolated occurrence of the so-called “Olenus“ beds of Mt. Jubilee. Japanese Journal of Geology and Geography, 15:149192.Google Scholar
Kobayashi, T. 1955. The Ordovician fossils from the McKay Group in British Columbia, western Canada, with a note on the early Ordovician paleogeography. Journal of Faculty of Science, University of Tokyo, Section 2, 9:355493.Google Scholar
Kurtz, V. E. 1975. Franconian (upper Cambrian) faunas from the Elvins Group of southeast Missouri. Journal of Paleontology, 49:10091043.Google Scholar
Lenz, A. C., Jin, J., McCracken, A. D., Utting, J., and Westrop, S. R. 1993. Paleoscene 15. Paleozoic biostratigraphy. Geoscience Canada, 20:4173.Google Scholar
Linarsson, J. G. O. 1869. Om Vestergötlands Cambriska och Siluriska aflagringar. Kungliga Svenska Vetenskapsakademiens Handlingar, 8, 89 p.Google Scholar
Lisogor, K. A. 1970. New genera of trilobites from the Upper Cambrian of Malyi Karatau. Geologia, 6:1320.Google Scholar
Loch, J. D., and Taylor, J. E. 1995. High-resolution biostratigraphy in the Upper Cambrian Ore Hill Member of the Gatesburg Formation, south-central Pennsylvania. In Mann, K. O. and Lane, H. R. (eds.), Graphic Correlation. Society for Sedimentary Geology. Special Publication, 53:131137.CrossRefGoogle Scholar
Lochman, C. 1958. Sulcocephalus Wilson, 1948, to replace Berkeia Resser, 1937, and Burnetiella to replace Burnetia Walcott, 1924. Journal of Paleontology, 32:247.Google Scholar
Lochman, C. 1964. Upper Cambrian faunas from the subsurface Deadwood Formation, Williston Basin, Montana. Journal of Paleontology, 38:3360.Google Scholar
Lu, Y. H., Chang, W. T., Chu, C. L., Chien, Y. Y., and Hsiang, L. W. 1965. Chinese Fossils of all Groups, Trilobites of China 1:1-362; 2:363-766. Science Press, Beijing. (In Chinese.).Google Scholar
Lu, Y. H., Chu, C. L., Chien, Y. Y., Lin, H. L., Zhou, Z. Y., and Yuan, K. X. 1974. Bio-environmental control hypothesis and its application to the Cambrian biostratigraphy and palaeozoogeography. Memoirs of the Nanjing Institute of Geology and Palaeontology, Academia Sinica 5:27110. (In Chinese.).Google Scholar
Lu, Y. H., and Lin, H. L. 1980. Cambro-Ordovician boundary in western Zhejiang and the trilobites contained therein. Acta Palaeontologica Sinica, 19:118134.Google Scholar
Lu, Y. H., and Lin, H. L. 1983. Uppermost Cambrian and lowermost Ordovician trilobites of the Jiangshan-Changshan area. In Nanjing Institute of Geology and Paleontology (ed.), Papers for the Symposium on the Cambrian-Ordovician and Ordovician-Silurian boundaries, Nanjing, 1983:611.Google Scholar
Lu, Y. H., and Lin, H. L. 1984. Late Late Cambrian and earliest Ordovician trilobites of Jiangshan-Changshan area, Zhejiang. In Stratigraphy and Paleontology of Systemic Boundaries in China. Cambrian-Ordovician Boundary, Volume 1:45143. Anhui Science and Technology Publishing House, Nanjing.Google Scholar
Lu, Y. H., and Lin, H. L. 1989. The Cambrian trilobites of western Zhejiang. Palaeontologia Sinica, New Series B, 25, 287 p. (In Chinese with English summary.).Google Scholar
Ludvigsen, R. 1979. Fossils of Ontario, Part 1. The trilobites. Royal Ontario Museum Life Sciences, Miscellanceous Publications, 96 p.Google Scholar
Ludvigsen, R. 1982. Upper Cambrian and Lower Ordovician trilobite biostratigraphy of the Rabbitkettle Formation, western District of Mackenzie. Royal Ontario Museum Life Sciences Contributions 134, 188 p.Google Scholar
Ludvigsen, R. 1991. The median suture of Asaphida: a monophyletic character? The Trilobite Papers, 3:1113.Google Scholar
Ludvigsen, R. 1992. The search for the oldest fossils in the Cordillera. In Levson, V. (compiler), Pioneering Geology in the Canadian Cordillera. British Columbia Geological Survey Branch Open File 1992-19:6771.Google Scholar
Ludvigsen, R., and Pugh, K. W. 1995. A trilobite from an archeological site along the Fraser River. Abstract, First British Columbia Paleontological Symposium, Courtenay, British Columbia:7.Google Scholar
Ludvigsen, R., and Tuffnell, P. 1994. The last olenacean trilobite: Triarthrus in the Whitby Formation (Upper Ordovician) of southern Ontario. In Landing, E. (ed.), Studies on Stratigraphy and Paleontology in Honor of Donald W. Fisher. New York State Museum Bulletin, 481:183212.Google Scholar
Ludvigsen, R., and Westrop, S. R. 1983a. Franconian trilobites of New York State. New York State Museum and Science Service Memoir, 23, 83 p.Google Scholar
Ludvigsen, R., and Westrop, S. R. 1983b. Trilobite biofacies of the Cambrian-Ordovician boundary interval in northern North America. Alcheringa, 7:301319.Google Scholar
Ludvigsen, R., and Westrop, S. R. 1985. Three new Upper Cambrian stages for North America. Geology, 13:139143.2.0.CO;2>CrossRefGoogle Scholar
Ludvigsen, R., Westrop., S. R., and Kindle, C. H. 1989. Sunwaptan (Upper Cambrian) trilobites of the Cow Head Group, western Newfoundland, Canada. Palaeontographica Canadiana, 6, 175 p.Google Scholar
Ludvigsen, R., Westrop., S. R., Pratt, B. R., Tuffnell, P. A., and Young, G. A. 1986. Palaeoscene 3. Dual biostratigraphy: zones and biofacies. Geoscience Canada, 13:139154.Google Scholar
McNamara, K. J. 1986. The role of heterochrony in the evolution of Cambrian trilobites. Biological Reviews, 61:121156.Google Scholar
M'Coy, F. 1849. On the classification of some British fossil Crustacea, with notices of new forms in the University collection at Cambridge. Annual Magazine of Natural History (London), Series 2, 4:161179, 392-414.Google Scholar
Moore, R. C. (ed.). 1959. Treatise on invertebrate paleontology, Part O, Arthropoda I. Geological Society of America and University of Kansas Press, 560 p.Google Scholar
Mott, J. A. 1989. Structural and stratigraphic relations in the White River region, eastern Main Ranges, southern Canadian Rocky Mountains, British Columbia. , Queen's University, Kingston, Ontario, 404 p.Google Scholar
Navás, J. L. 1936. Schwedisch-chinesisch wissenschfliche Expedition nach den nordwestlichen Provinzen Chinas unter Leitung von Dr Sven Hedin und Prof. Sü Ping-chan, Insekten, 29, Plecoptera. Arkiv für Zoologie, 27A, 1.Google Scholar
Nikolaisen, F., and Henningsmoen, G. 1985. Upper Cambrian and lower Tremadoc olenid trilobites from the Digermul peninsula, Finnmark, northern Norway. Norges geologiske undersøkelse, Bulletin 400, 49 p.Google Scholar
Norford, B. S. 1962. Illustrations of Canadian fossils. Cambrian, Ordovician and Silurian of the western Cordillera. Geological Survey of Canada Paper 62-14, 25 p.Google Scholar
Öpik, A. A. 1963. Early Upper Cambrian fossils from Queensland. Bureau of Mineral Resources, Geology and Geophysics of Australia, Bulletin 64, 133 p.Google Scholar
Öpik, A. A. 1967. The Mindyallan fauna of northwestern Queensland. Bureau of Mineral Resources, Geology and Geophysics of Australia, Bulletin 74, 2 volumes, 404 p., 167 p. Google Scholar
Palmer, A. R. 1954. The faunas of the Riley Formation in central Texas. Journal of Paleontology, 28:709786.Google Scholar
Palmer, A. R. 1960a. Trilobites of the Upper Cambrian Dunderberg Shale. Eureka District. Nevada. United States Geological Survey Professional Paper 334c, 109 p.Google Scholar
Palmer, A. R. 1960b. The type species of Litocephalus Resser, 1937 (Trilobita). Journal of Paleontology, 30:608610.Google Scholar
Palmer, A. R. 1962. Glyptagnostus and associated trilobites in the United States. United States Geological Survey Professional Paper, 374F, 49 p.Google Scholar
Palmer, A. R. 1965. Trilobites of the Late Cambrian Pterocephaliid Biomere in the Great Basin, United States. United States Geological Survey Professional Paper 493, 105 p.CrossRefGoogle Scholar
Palmer, A. R. 1968. Cambrian trilobites of east-central Alaska. Lower Paleozoic paleontology and stratigraphy of east-central Alaska. United States Geological Survey Professional Paper 559-B, 115 p.Google Scholar
Peng, S. 1990. Upper Cambrian of the Cili-Taoyuan area, Hunan and its trilobite succession. Journal of Stratigraphy, 14:261276. (In Chinese.).Google Scholar
Peng, S. 1992. Upper Cambrian biostratigraphy and trilobite faunas of the Cili-Taoyuan area, northwestern Hunan, China. Memoirs of the Australasian Association of Palaeontologists, 13, 119 p.Google Scholar
Pratt, B. R. 1992. Trilobites of the Marjuman and Steptoean stages (Upper Cambrian), Rabbitkettle Formation, southern Mackenzie Mountains, northwest Canada. Palaeontographica Canadiana 9, 179 p.Google Scholar
Rasetti, F. 1951. Middle Cambrian stratigraphy and faunas of the Canadian Rocky Mountains. Smithsonian Institution Miscellaneous Contributions, 116, 277 p.Google Scholar
Resser, C. E. 1938. Fourth contribution to nomenclature of Cambrian trilobites. Smithsonian Miscellaneous Collections, 97, 43 p.Google Scholar
Resser, C. E. 1942. New Upper Cambrian trilobites. Smithsonian Miscellaneous Collections, 103:1136.Google Scholar
Robison, R. A. 1964. Late Middle Cambrian faunas from western Utah. Journal of Paleontology, 38:510566.Google Scholar
Robison, R. A. 1971. Additional Middle Cambrian trilobites from the Wheeler Shale of Utah. Journal of Paleontology, 45:796804.Google Scholar
Robison, R. A. 1988. Trilobites of the Holm Dal Formation (late Middle Cambrian) of central North Greenland. Meddelelser om Grønland. Geoscience, 20:20103.CrossRefGoogle Scholar
Roemer, F. 1849. Texas, mit besonderer Rucksicht auf deutsche Auswanderung und die physichen Verhaltnisse des Landes, 14. Bonn, Germany, 464 p.Google Scholar
Rudkin, D. M. 1979. Healed injuries in Ogygopsis klotzi (Trilobita) from the Middle Cambrian of British Columbia. Royal Ontario Museum, Life Sciences Occasional Paper 32, 8 p.Google Scholar
Rudkin, D. M. 1996. The Trilobite Beds of Mount Stephen, Yoho National Park, p. 5968. In Ludvigsen, R. (ed.), Life in Stone: A Natural History of British Columbia's Fossils. University of British Columbia Press, Vancouver.Google Scholar
Rushton, A. W. A. 1967. The Upper Cambrian trilobite Irvingella nuneatonensis (Sharman). Palaeontology, 10:339348.Google Scholar
Rushton, A. W. A. 1983. Trilobites from the Upper Cambrian Olenus Zone in central England. Special Papers in Palaeontology, 30:107139.Google Scholar
Salter, J. W. 1864. A monograph of the British trilobites. Palaeontographical Society (London) Monograph, p. 81128.Google Scholar
Sharman, G. A. 1886. On the new species Olenus nuneatonensis and Obolella granulata, from the Lower Silurian (‘Cambrian’, Lapworth), near Nuneaton. Geological Magazine, 3:565566.Google Scholar
Shaw, A. B. 1951. The paleontology of northwestern Vermont. I. New Late Cambrian trilobites. Journal of Paleontology, 25:97114.Google Scholar
Shergold, J. H. 1972 Late Upper Cambrian trilobites from the Gola Beds, western Queensland. Bureau of Mineral Resources, Geology and Geophysics of Australia, Bulletin 112:189.Google Scholar
Shergold, J. H. 1975. Late Cambrian and Early Ordovician trilobites of the Burke River structural belt, western Queensland. Bureau of Mineral Resources, Geology and Geophysics of Australia, Bulletin 153:1251.Google Scholar
Shergold, J. H. 1977. Classification of the trilobite Pseudagnostus . Palaeontology, 20:69100.Google Scholar
Shu, D., Geyer, G., Ling, C., and Xingliang, Z. 1995. Redlichacean trilobites with preserved soft parts from the Lower Cambrian Chengjiang fauna (South China). Beringeria Special Issue 2. Morocco 1995. The Lower and Middle Cambrian standard of western Gondwana. 203240.Google Scholar
Shumard, B. F. 1861. The primordial zone of Texas with descriptions of new fossils. American Journal of Science, s. 2, 32:213221.Google Scholar
Speyer, S. E. 1987. Comparative taphonomy and paleoecology of trilobite lagerstätten. Alcheringa, 11:205232.Google Scholar
Speyer, S. E., and Brett, C. E. 1985. Clustered trilobite assemblages in the Middle Devonian Hamilton Group. Lethaia, 18:85103.Google Scholar
Speyer, S. E., and Brett, C. E. 1986. Trilobite taphonomy and Middle Devonian taphofacies. Palaois, 1:312327.Google Scholar
Stitt, J. H. 1971. Late Cambrian and earliest Ordovician trilobites, Timbered Hills and Lower Arbuckle groups, western Arbuckle Mountains, Murray County, Oklahoma. Oklahoma Geological Survey Bulletin, 110, 83 p.Google Scholar
Stitt, J. H. 1983. Enrolled Late Cambrian trilobites from the Davis Formation, southeast Missouri. Journal of Paleontology, 57:93105.Google Scholar
Taylor, M. E. 1976. Indigenous and redeposited trilobites from Late Cambrian basinal environments of central Nevada. Journal of Paleontology, 50:668700.Google Scholar
Taylor, M. E., and Robison, R. A. 1976. Trilobites in Utah folklore. Brigham Young University Geology Studies, 23:15.Google Scholar
Troedsson, G. T. 1937. On the Cambro-Ordovician faunas of western Quruq Tagh, eastern T'ien Shan. Paleontologica Sinica, New Series B, number 2, whole series number 106, 74 p.Google Scholar
Troedsson, G. T. 1951. Hedinaspis, new name for Hedinia Troedsson, non Navás. Geologiska föreningens i Stockholm Förhandlingar, 73:695.Google Scholar
Tullberg, S. A. 1880. Om Agnostus-aterna i de Kambriska aflagringarna vid Andrarum. Sveriges Geologiska Undersökning, Series C. 42, 37 p.Google Scholar
Walcott, C. D. 1908. Mount Stephen rocks and fossils. Canadian Alpine Journal, 1:232248.Google Scholar
Walcott, C. D. 1924. Cambrian geology and paleontology, 5, no. 3. Cambrian and lower Ozarkian trilobites. Smithsonian Miscellaneous Collections, 75:5360.Google Scholar
Wang, J. (ed.). 1964. Handbook of Index Fossils of China, Hunan Region. Geology Press, Beijing, 173 p. (In Chinese.).Google Scholar
Webby, B. D., Wang, Q., and Mills, K. J. 1988. Upper Cambrian and basal Ordovician trilobites from western New South Wales. Palaeontology, 31:905938.Google Scholar
Westrop, S. R. 1986. Trilobites of the Upper Cambrian Sunwaptan Stage, southern Canadian Rocky Mountains, Alberta. Palaeontographica Canadiana, 3, 179 p.Google Scholar
Westrop, S. R. 1989. Facies anatomy of an Upper Cambrian grand cycle: Bison Creek and Mistaya formations, southern Alberta. Canadian Journal of Earth Sciences, 26:22922304.Google Scholar
Westrop, S. R. 1995. Sunwaptan and Ibexian (Upper Cambrian-Lower Ordovician) trilobites of the Rabbitkettle Formation, Mountain River region, northern Mackenzie Mountains, northwest Canada. Palaeontographica Canadiana, 12, 75 p.Google Scholar
Whitehouse, F. W. 1936. The Cambrian faunas of northeastern Australia, Parts 1-2. Memoirs of the Queensland Museum, 11:59112.Google Scholar
Whitehouse, F. W. 1939. The Cambrian faunas of northeastern Queensland. Part 3. The polymerid trilobites. Memoirs of the Queensland Museum, 21:179282.Google Scholar
Whittington, H. B. 1980. Exoskeleton, moult stage, appendage morphology, and habits of the Middle Cambrian trilobite Olenoides serratus . Palaeontology, 23:171204.Google Scholar
Whittington, H. B. 1990. Articulation and exuviation in Cambrian trilobites. Philosophical Transactions of the Royal Society of London: Series B, Biological Sciences, 329:2746.Google Scholar
Whittington, H. B. 1992. Fossils Illustrated Volume 2. Trilobites. Boydell Press, Woodbridge, Suffolk, 145 p.Google Scholar
Wilson, J. L. 1949. The trilobite fauna of the Elvinia Zone in the basal Wilberns Limestone of Texas. Journal of Paleontology, 23:2544.Google Scholar
Wilson, J. L. 1951. Franconian trilobites of the central Appalachians. Journal of Paleontology, 25:617654.Google Scholar
Wilson, J. L. 1954. Late Cambrian and Early Ordovician trilobites from the Marathon Uplift, Texas. Journal of Paleontology, 28:249286.Google Scholar
Wilson, J. L., and Frederickson, E. A. 1950. The Irvingella major (“Ptychopleurites“) faunizone of the Upper Cambrian. American Journal of Science, 248:891902.Google Scholar
Xiang, L. W., and Zhang, T. R. 1985. Description of trilobites. Ministry of Geology and Mineral Resources, Geology Memoir Series 2, 4:64243.Google Scholar
Young, G. A., and Ludvigsen, R. 1989. Mid-Cambrian trilobites from the lowest part of the Cow Head Group, western Newfoundland. Geological Survey of Canada, Bulletin 392, 49 p.Google Scholar