Skip to main content Accessibility help

Paracalamitina striata—a newly reconstructed equisetophyte from the Permian of Angaraland

  • Serge V. Naugolnykh (a1)


This paper deals with Paracalamitina striata (Schmalhausen) Zalessky, 1934, an equisetophyte of the family Tschernoviaceae from the Permian of Angaraland. New material contributes to a better understanding of this plant, which now can be reconstructed. Paracalamitina striata is an equisetopsid with opposite ribs on the stem. Its main stem bears lateral shoots or shoot and leaf scars, that are located at nodes. Shoot nodes have subcylindrical leaf sheaths. The distal parts of leaves are free and acute. Generative organs are fertile zones, which are formed by numerous sporangiophores with peltate sporophylls. A terminal sterile whorl is presented above each fertile zone. An emended diagnosis of Paracalamitina Zalessky is provided on the basis of new material and re-examination of previously collected specimens as well. The nomenclature and taxonomic status of P. striata also are discussed.



Hide All
Anderson, J. M., and Anderson, H. M. 1985. Palaeoflora of Southern Africa, Prodromus of South African Megafloras; Devonian to Lower Cretaceous. A.A. Balkema, Rotterdam, 432.
Barthel, M. 1980. Calamiten aus dem Oberkarbon und Rotliegenden des Thuringer Waldes. Museum fur Naturkunde, 100 Jahre Arboretum, p. 237257.
Barthel, M., and Rossler, R. 1996. Palaeontologische Fundschichten in Rotliegend von Manebach (Thuringer Wald) mit Calamites gigas (Sphenophyta). Veroffentlichungen Naturhistorische Museum Schleusingen, 11:321.
Chandra, S. 1992. Changing patterns of the Permian Gondwana vegetation. Palaeobotanist, 40:73100.
Cichan, M. A. 1986. Vascular cambium and wood development in Carboniferous plants, III, Arthropitys (Equisetales; Calamitaceae). Canadian Journal of Botany, 64:688695.
Cichan, M. A., and Taylor, T. N. 1983. A systematic and developmental analysis of Arthropitys deltoides p. nov. Botanical Gazette, 144:285294.
Duran, M., Hunicken, M. A., and Anton, A. M. 1997. Novedos hallazgos de Sphenopsida en la Formacion Bajo de Veliz, provincia de San Luis, Argentina. Ameghiniana (Review of Association of Paleontology, Argentina), 34(3):259264.
Gastaldo, R. A. 1981. Palaeostachya dircei n. sp. an autigenically cemented equisetalean strobilus from the Middle Pennsylvanian of southern Illinois. American Journal of Botany, 68:13061318.
Gorelova, S. C., Menshikova, L. V., and Khalfin, L. L. 1973. Phytostratigraphy and Identification of Plants from late Palaeozoic Coal Deposits in the Kuznetsk Basin. Kemerovo's Knizhnoe Izdatelstvo, Kemerovo, 168 p.
Hirmer, M. 1927. Handbuch der Palaobotanik, Bd 1. R. Oldenbourg, Berlin, 708 p.
Kerp, H. 1984. Aspects of Permian Palaeobotany and Palynology, V, On the nature of Asterophyllites dumasii Zeiller, its correlation with Catamites gigas Brongniart and the problem concerning its sterile foliage. Review of Palaeobotany and Palynology, 41:301317.
Langiaux, J., and Marguerier, J. 1980. Paleoxylologie du bassin Stephanien de Blanzy, 2, Decouverte d'une structure d'Arthropytis communis, considerations taxonomiques et phylogenetigues. La physiophyle, 92:89102.
Mamay, S. H., and Bateman, R. M. 1991. Archaeocalamites lazarii, sp. nov.: the range of Archaeocalamitaceae extended from the lowermost Pennsylvanian to the mid-Lower Permian. American Journal of Botany, 78:489496.
Mcloughlin, S. 1992. Permian sphenophytes from the Collie and Perth Basins, Western Australia. Review of Palaeobotany and Palynology, 75:153182.
Meyen, S. V. 1971. Phyllotheca-like plants from the Upper Palaeozoic flora of Angaraland // Palaeontographica, Abt B, 133:133.
Meyen, S. V. 1982. The Carboniferous and Permian floras of Angaraland (a synthesis). Biological Memoirs, 7:1110.
Meyen, S. V. 1987. Fundamentals of Palaeobotany. Chapman and Hall, London, 432 p.
Meyen, S. V., and Menshikova, L. V. 1983. Systematics of the Upper Palaeozoic articulates of the family Tchernoviaceae. Botanical Journal (St-Petersburg), 68:721729. (In Russian)
Naugolnykh, S. V. 1998a. Comparative analysis of the Permian floristic assemblages of Kozhim section (the Pechora Pre-Urals region) and the stratotype section (Middle Urals region), The Urals: fundamental problems of geodynamics and stratigraphy. Nauka, Moscow, 154182.
Naugolnykh, S. V. 1998b. Kungurian flora of the Middle Cis-Urals. Geos, Moscow, 201 p.
Neuburg, M. F. 1964. Permian flora of Petchora basin, Pt. II, Sphenopsida. Nauka, Moscow, 92 p. (In Russian)
Pant, D. D., Nautiyal, D. D., and Misra, L. 1981. Giridia indica gen. et sp. nov., the possible cone of Phyllotheca indica Bunbury. Palaeonthographica, 176(5–6):174178.
Rasskasova, E. S. 1961. Sphenophytes from the Upper Paleozoic of the Tunguska basin, Pt. 1–2. Annals of the Scientific Institute of Arctic Geology, 23–24:173.
Rigby, J. F. 1989. Austroannularia gen. nov., an asymmetrical Permian sphenopsid leaf whorl from Gondwanaland. Palaeobotanist, 37(2):189191.
Schmalhausen, I. F. 1879. Beitrage zur Jura-Flora Russlands.— Memoirs of Academy of Sciences, St. Petersburg, 27(4):196.
Schmalhausen, I. F. 1881. Jurassic Flora of the Kuznetsk basin and the Pechora area. Transactions of St. Petersburg Mineralogical Society, 16:97178.
Taylor, T. N., and Taylor, E. L. 1993. The Biology and Evolution of Fossil Plants. Prentice Hall, Inc., New Jersey, 982 p.
Tidwell, W. D., Jennings, J. R., and Call, V. B. 1988. Flora of Manning Canyon Shale, Pt. III, Sphenophyta. Brigham Young Univrsity, Geology Studies, 35:1532.
Zalessky, M. D. 1927. Flore permienne des limites ouraliennes d'Angaride. Atlas. Memoirs, Commite Geologique. St. Petersburg, 176:552.
Zalessky, M. D. 1934. Observations sur les vegetaux permiens du bassin de la Petchora, I. Bulletin of Academy of Sciences, URSS, Mathematics and Natural Sciences, 2–3:241290.

Related content

Powered by UNSILO

Paracalamitina striata—a newly reconstructed equisetophyte from the Permian of Angaraland

  • Serge V. Naugolnykh (a1)


Full text views

Total number of HTML views: 0
Total number of PDF views: 0 *
Loading metrics...

Abstract views

Total abstract views: 0 *
Loading metrics...

* Views captured on Cambridge Core between <date>. This data will be updated every 24 hours.

Usage data cannot currently be displayed.