Hostname: page-component-76fb5796d-dfsvx Total loading time: 0 Render date: 2024-04-26T10:32:33.251Z Has data issue: false hasContentIssue false

A new Ladinian (Middle Triassic) mysidacean shrimp (Crustacea, Lophogastrida) from northern Italy and southern Switzerland

Published online by Cambridge University Press:  22 October 2019

Cristiano Larghi
Affiliation:
Eni Spa – Exploration, 5th Office Building, Via Emilia 1, 20097, San Donato Milanese, Italy
Andrea Tintori
Affiliation:
Dipartimento di Scienze della Terra ‘Ardito Desio’, Università degli Studi di Milano, via Mangiagalli 34, I-20133Milano, Italy. A.T.
Daniela Basso
Affiliation:
Dipartimento di Scienze dell'Ambiente e della Terra, Università degli Studi di Milano-Bicocca, Piazza della Scienza 4, 20126Milano, Italy
Gianluca Danini
Affiliation:
Civico Museo Insubrico di Storia Naturale di Clivio/Induno Olona
Markus Felber
Affiliation:
CGA SAGL, Casella Postale 124 6834-Morbio Inferiore, Switzerland MCSN, Lugano, Switzerland

Abstract

Since the end of the nineteenth century, many paleontological studies have been developed on the Middle Triassic localities of the Monte San Giorgio area (Canton Ticino, Switzerland, and Lombardy, Italy). These localities were inscribed in 2003 and 2010 in the UNESCO World Heritage List due to their paleontological relevance. New crustaceans are here described from the Kalkschieferzone member of the upper Ladinian Meride Limestone, outcropping near Besnasca-Cà del Frate (Viggiù, Italy) and Meride (Mendrisio, Switzerland). Several hundred specimens of a new genus and new species of Lophogastrida, named here Vicluvia lombardoae, improve information on Triassic mysidaceans (sensu lato), a group of shrimp-like crustaceans presently including hundreds of species worldwide distributed in marine and freshwater environments. The paper discusses the relations between Vicluvia n. gen. and the other fossil genera belonging to the same family. Vicluvia lombardoae n. gen. n. sp. exhibits two dorsally vanishing transverse grooves with their branches regularly bending toward the posterior margin of the shield and a telson with a rounded and setose apex. It was probably a euryhaline species living in a transitional environment affected by frequent and ephemeral salinity variations. Middle Triassic taphonomic windows have preserved specimens of lophogastrids in shallow basins along the Tethys, from the present Alps to China as well as in the German Basin.

UUID: http://zoobank.org/1623e8f9-2ceb-4cd6-bcbe-4f05a873e1ff

Type
Articles
Copyright
Copyright © 2019, The Paleontological Society

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

Footnotes

3

Current address: TRIASSICA – Institute for Triassic Fossil Lagerstätten, 23828 Perledo (LC), Italy

References

Affer, D., and Teruzzi, G., 1999, Thylacocephalan crustaceans from the Besano Formation, Middle Triassic, N. Italy: Rivista del Museo civico ‘E. Caffi’ di Bergamo, v. 20, p. 58.Google Scholar
Azeiteiro, U.M.M., Jesus, L., and Marques, J.C., 1999, Distribution, population dynamics and production of the suprabenthic mysid Mesopodopsis slabberi Van Beneden in the Mondego estuary (Western coast of Portugal): Journal of Crustacean Biology, v. 19, p. 498509.10.2307/1549259CrossRefGoogle Scholar
Azeiteiro, U.M., Fonseca, J., and Marques, J.C.S., 2001, Biometry, estimates of production and seasonal variation in the biochemical composition of Mesopodopsis slabberi (Van Beneden, 1861) (Crustacea: Mysidacea): Boletín Instituto Español de Oceanografía, v. 17, p. 1525.Google Scholar
Bechly, G., and Stockar, R., 2011, The first Mesozoic record of the extinct apterygote insect genus Dasyleptus (Insecta: Archaeognatha: Monura: Dasyleptidae) from the Triassic of Monte San Giorgio (Switzerland): Palaeodiversity, v. 4, p. 2337.Google Scholar
Benton, M.J., 1993, The Fossil Record 2: London, Chapman and Hall, 845 p.Google Scholar
Bernasconi, S., and Riva, A., 1993, Organic geochemistry and depositional environment of a hydrocarbon source rock: The Middle Triassic Grenzbitumnzone Formation, southern Alps, Italy/Switzerland, in Spencer, A.M., ed., Generation, Accumulation and Production of Europe's Hydrocarbons III: Berlin, Springer-Verlag, 380 p.Google Scholar
Beurlen, K., 1930, Vergleichende Stammgeschichte Grundlagen Methoden. Probleme unter besondere Berucksichtigung dcr hoheren Krebse: Forschung der Geologie und Paleontologie, v. 8, p. 317586.Google Scholar
Bill, P.C., 1914, Über Crustaceen aus dem Voltziensandstein des Elsasses: Mitteilungen der Geologischen Landesanstalt Elsass-Lothringen, v. 8, p. 289338.Google Scholar
Boas, J.E.V., 1883, Studien uber die Verwandtschaftsbeziehungen der Malakostraken: Morphologisches Jahrbuch, v. 8, p. 485579.Google Scholar
Broili, F., 1917, Eine neue Crustaceen Form aus dem lithographischen Schiefer des oberen Jura von Franken: Zentralblatt für Mineralogie, Geologie und Paläontologie Jahrbuch, v. 1917, p. 426429.Google Scholar
Brusca, R.C., and Brusca, G.J., 1990, Invertebrates: Sunderland, Massachusetts, Sinauer Associates, 922 p.Google Scholar
Bürgin, T., Eichenberger, U., Furrer, H., and Schanz, T.K., 1991, Die Prosanto-Formation eine fischreiche Fossil-Lagerstätte in der Mitteltrias der Silvretta-Decke (Kanton Graubünden, Schweiz): Eclogae Geologicae Helvetiae, v. 84, no. 3, p. 921990.Google Scholar
Dana, J.D., 1852, Crustacea, part I, in United States Exploring Expedition, During the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U.S.N, Vol. 13: Philadelphia, C. Sherman, 692 p.Google Scholar
De Alessandri, G., 1910, Studii sui pesci triasici della Lombardia: Memorie della Societa` Italiana di Science Naturali, v. 7, p. 1147.Google Scholar
Etter, W., 1994, A new penaeid shrimp (Antrimpos mirigiolensis n. sp., Crustacea, Decapoda) from the Middle Triassic of the Monte San Giorgio (Ticino, Switzerland): Neues Jahrbuch fur Geologie und Palaontologie Monatshefte, v. 4, p. 223230.10.1127/njgpm/1994/1994/223CrossRefGoogle Scholar
Feldmann, R.M., Schweitzer, C.E., Hu, S., Zhang, Q., Zhou, C., Xie, T., Huang, J., and Wen, W., 2012, Macrurous Decapoda from the Luoping Biota (Middle Triassic) of China: Journal of Paleontology, v. 86, p. 425441.10.1666/11-113.1CrossRefGoogle Scholar
Feldmann, R.M., Schweitzer, C.E., Hu, S., Huang, J., Zhou, C., Zhang, Q., Wen, W., Xie, T., and Maguire, E., 2016, Spatial distribution of Crustacea and associated organisms in the Luoping biota (Anisian, Middle Triassic), Yunnan Province, China: Evidence of periodic mass kills: Journal of Paleontology, doi:10.1017/jpa.2015.60.Google Scholar
Feldmann, R.M., Schweitzer, C.E., Hu, S., Huang, J., Zhou, C., Zhang, Q., Wen, W., Xie, T., Schram, F.R., and Jones, W.T., 2017, Earliest occurrence of lophogastrid mysidacean arthropods (Crustacea, Eucopiidae) from the Anisian Luoping Biota, Yunnan Province, China: Journal of Paleontology, v. 91, p. 100115.CrossRefGoogle Scholar
Frauenfelder, A., 1916, Beiträge zur Geologie der Tessiner Kalkalpen: Eclogae geologicae Helvetiae, v. 14, p. 247367.Google Scholar
Fu, W.L., Wilson, G.D.F., Jiang, D.Y., Sun, Y.L., Hao, W.C., and Sun, Z.Y., 2010, A new species of Protamphisopus Nicholls (Crustacea, Isopoda, Phreatoicidea) from Middle Triassic Luoping Fauna of Yunnan Province: China: Journal of Paleontology, v. 84, p. 10031013.10.1666/10-042.1CrossRefGoogle Scholar
Furrer, H., 1995, The Kalkschieferzone (Upper Meride Limestone; Ladinian) near Meride (Canton Ticino, southern Switzerland) and the evolution of a Middle Triassic intraplatform basin: Eclogae geologicae Helvetiae, v. 88, p. 827852.Google Scholar
Gall, J.C., 1971, Faunes et paysages du Grès à Voltzia du Nord des Vosges: Essai paléoécologique sur le buntsandstein supérieur, Université de Strasbourg, Service de la carte géologique d'Alsace et de Lorraine, Mémoires, v. 34, p. 1318.Google Scholar
Gall, J.C., and Grauvogel-Stamm, L., 2005, The early Middle Triassic ‘Grès à Voltzia’ Formation of eastern France: A model of environmental refugium: Comptes Rendus Palevol, v. 4, p. 637652.10.1016/j.crpv.2005.04.007CrossRefGoogle Scholar
Gozzi, E., Lombardo, C., and, Larghi, C., 2003, Exceptional fossil preservation in the Besano-Monte San Giorgio Area: The case of the arthropods: Abstract and Poster, EPA Workshop 2003, Exceptional Preservation, Teruel, Spain, 24–27 September, p. 5961.Google Scholar
Greenwood, J.G., Jones, M.B., and Greenwood, J., 1989, Salinity effects on brood maturation of the mysid crustacean Mesopodopsis slabberi: Marine Biological Association (United Kingdom) Journal, v. 69, p. 683694.10.1017/S0025315400031064CrossRefGoogle Scholar
Haworth, A.H., 1825, A new binary arrangement of the macrurous Crustacea: Philosophical Magazine and Journal of Natural Philosophy, Chemistry and Arts, v. 65, p. 183184.Google Scholar
Hessler, R.R., 1969, Peracarida, in Moore, R.C., ed., Treatise on Invertebrate Paleontology, Part R, Arthropoda 4, Volume 1: Boulder, Colorado, and Lawrence, Kansas, Geological Society of America and University of Kansas Press, p. R360R393.Google Scholar
Ji, C., Sang, W.L., Diao, G.W., Motani, R., and Tintori, A., 2012, Biodiversity and stratigraphic distribution of the first late Ladinian (Middle Triassic) marine vertebrate fauna (Xingyi Fauna) from South China: Journal of Vertebrate Paleontology, v. 32, 131 p.Google Scholar
Jiang, D., Motani, R., Hao, W., Rieppel, O., Sun, Y., Tintori, A., Sun, Z., and Schmitz, L., 2009, Biodiversity and sequence of the Middle Triassic Panxian marine reptile fauna, Guizhou Province, China: Acta Geologica Sinica, v. 83, p. 451459.CrossRefGoogle Scholar
Križnar, M., and Hitij, T., 2010, Nevretenčarji (Invertebrates) Strelovške formacij: Scopolia Supplementum. v. 5, p. 91107.Google Scholar
Krzeminski, W., and Lombardo, C., 2001, New fossil Ephemeroptera and Coleoptera from the Ladinian (Middle Triassic) of Canton Ticino (Switzerland): Rivista Italiana di Paleontologia e Stratigrafia, v. 107, p. 6978.Google Scholar
Larghi, C., and Tintori, A., 2007, First record of a decapod from the Meride Limestone: New data from one of the best Ladinian (Middle Triassic) taphonomic windows of a transitional environment: Memorie della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, v. 35, p. 6869.Google Scholar
Latreille, P.A., 1802, Histoire naturelle générale et particulière des Crustacés et des insectes. Tome 3, Familles naturelles des genres: Paris, F. Dufart, 479 p.Google Scholar
Lombardo, C., 1999, Sexual dimorphism in a new species of the actinopterygian Peltopleurus from the Triassic of northern Italy: Palaeontology, v. 42, p. 741760.10.1111/1475-4983.00095CrossRefGoogle Scholar
Lombardo, C., 2001, Actynopterigians from the Middle Triassic of northern Italy and Canton Ticino (Switzerland): Anatomical descriptions and nomenclatural problems: Rivista Italiana di Paleontologia e Stratigrafia, v. 107, p. 345369.Google Scholar
Lombardo, C., Tintori, A., Felber, M., Danini, G., Moratto, D., Pacor, G., and Tentor, M., 1999, Faunal assemblages in the Kalschieferzone (Meride Limestone, Upper Ladinian) of Meride (Canton Ticino, Switzerland): Rivista del Museo Civico di Scienze Naturali ‘Enrico Caffi,’ v. 20 (supplement), p. 101104.Google Scholar
Lombardo, C., Tintori, A., and Tona, D., 2012, A new species of Sangiorgioichthys (Actinopterygii, Semionotiformes) from the Kalkschieferzone of Monte San Giorgio (Middle Triassic; Meride, Canton Ticino, Switzerland): Bollettino della Società Paleontologica Italiana, v. 51, no. 3, p. 203212.Google Scholar
Lu, H., Jiang, D.Y., Motani, R., Ni, P.G., Sun, Z.Y., Tintori, A., Xiao, S.Z., Zhou, M., Ji, C., and Fu, W.L., 2018, Middle Triassic Xingyi fauna: Showing turnover of marine reptiles from coastal to oceanic environments: Palaeoworld, v. 27, p. 17116.CrossRefGoogle Scholar
Martin, J.W., and Davis, G.E., 2001, An Updated Classification of the Recent Crustacea: Los Angeles, Natural History Museum of Los Angeles County, 124 p.Google Scholar
Montagna, M., Haug, J.T., Strada, L., Haug, C., Felber, M., and Tintori, A. 2017, Central nervous system and muscular bundles preserved in a 240 million year old giant bristletail (Archaeognatha, Machilidae): Science Reports, v. 7, 46016, doi:10.1038/srep46016.Google Scholar
Motani, R., Jiang, D.Y., Tintori, A., Sun, Y.L., Hao, W.C., Boyd, A., Hinic-Frlog, S., Schmitz, L., Shin, J.Y., and Sun, Z.Y., 2008, Horizons and assemblages of Middle Triassic marine reptiles from Panxian, Guizhou, China: Journal of Vertebrate Paleontology, v. 28, p. 900903.CrossRefGoogle Scholar
Münster, Von G., 1839, Decapoda Macroura. Abbildung und Beschreibung der fossilen langschwänzigen Krebse in den Kalkschiefern von Bayern: Beitrage Petrefactenkde, v. 2, p. 188.Google Scholar
Pinna, G., Arduini, P., Pesarini, C., and Teruzzi, G., 1982, Thylacocephala: una nuova classe di crostacei fossili: Atti della Società Italiana di Scienze Naturali e del Museo civico di Storia Naturale di Milano, v. 123, p. 469482.Google Scholar
Renesto, S., 1993, A juvenile Lariosaurus (Reptilia, Sauropterygia) from the Kalkschieferzone (Uppermost Ladinian) near Viggiù (Varese, northern Italy): Rivista Italiana di Paleontologia e Stratigrafia, v. 92, p. 199212.Google Scholar
Renesto, S., Lombardo, C., Tintori, A., and Danini, G., 2003, Nothosaurid embryos from the Middle Triassic of northern Italy: An insight into the viviparity of Nothosaurs: Journal Vertebrate Paleontology, v. 23, p. 958961.CrossRefGoogle Scholar
Riva, A., Salvatori, T., Cavaliere, R., Ricchiuto, T., and Novelli, L., 1985, Origin of oils in Po Basin, northern Italy: Organic Geochemistry, v. 10, p. 391400.CrossRefGoogle Scholar
Sars, G.O., 1885, Report on the Schizopoda collected by the HMS Challenger during the years 1873–1876, in Nartes, G.S., ed., Report on the Scientific Results of the Voyage of the H.M.S. Challenger during the Years 1873– 1876: London, Longmans & Co., p. 1–228, pls. 138.Google Scholar
Sars, M., 1857, Om 3 nye norske Krebsdyr: Forhandlimgar vid det Skandinaviska Naturforskaremotet, v. 7, p. 160175.Google Scholar
Scheuring, B.W., 1978, Mikrofloren aus den Meridenkalken des M.te San Giorgio (Kanton Tessin): Schweizerische Paläontologische Abhandlungen, v. 100, p. 1100.Google Scholar
Schram, F.R., 1984, Relationships within eumalacostracan crustaceans: Transactions of the San Diego Society of Natural History, v. 20, p. 301312.CrossRefGoogle Scholar
Schram, F.R., 1986, Crustacea: New York, Oxford University Press, 606 p.Google Scholar
Secretan, S., and Riou, B., 1986, Les Mysidaces (Crustacea, Peracarida) du Callovien de la Voulte-sur- Rhone: Annales de Paleontologie, v. 72, p. 295323.Google Scholar
Senn, A., 1924, Beiträge zur Geologie des Alpensüdrandes zwischen Mendrisio und Varese: Eclogae Geologicae Helvetiae, v. 18, p. 550632.Google Scholar
Stockar, R., 2010, Facies, depositional environment, and palaeoecology of the Middle Triassic Cassina beds (Meride Limestone, Monte San Giorgio, Switzerland): Swiss Journal of Geosciences, v. 103, p. 101119.10.1007/s00015-010-0008-2CrossRefGoogle Scholar
Stockar, R., and Garassino, A., 2013, Meridecaris ladinica n. gen. n. sp. (Crustacea, Decapoda, Clytiopsidae) from the Middle Triassic (Ladinian) of Monte San Giorgio (Canton Ticino, Switzerland): Neues Jahrbuch für Geologie und Palaontologie-Abhandlungen, v. 270, p. 347356.CrossRefGoogle Scholar
Stockar, R., and Kustatscher, E., 2010, The Ladinian flora from the Cassina beds (Meride limestone, Monte San Giorgio, Switzerland): Preliminary results: Rivista Italiana di Paleontologia e Stratigrafia, v. 116, p. 173188.Google Scholar
Stockar, R., Baumgartner, P.O., and Condon, D., 2012, Integrated Ladinian bio-chronostratigraphy and geochrononology of the Monte San Giorgio, (southern Alps, Switzerland): Swiss Journal of Geosciences, v. 105, p. 85108.CrossRefGoogle Scholar
Stockar, R., Adatte, T., Baumgartner, P.O., and Föllmi, K.B., 2013, Palaeoenvironmental significance of organic facies and stable isotope signatures: The Ladinian San Giorgio Dolomite and Meride Limestone of Monte San Giorgio (Switzerland, WHL UNESCO): Sedimentology, v. 60, p. 239269.CrossRefGoogle Scholar
Strada, L., Montagna, M., and Tintori, A., 2014, A new genus and species of the family Trachypachidae (Coleoptera, Adephaga) from the upper Ladinian (Middle Triassic) of Monte San Giorgio: Rivista Italiana di Paleontologia e Stratigrafia, v. 120, p. 183190.Google Scholar
Taylor, R.S., Schram, F.R., and Yan-Bin, S., 2001, A new upper Middle Triassic shrimp (Crustacea: Lophogastrida) from Guizhou, China, with discussion regarding other fossil ‘mysidaceans’: Journal of Paleontology, v. 75, p. 310318.CrossRefGoogle Scholar
Tintori, A., 1990a, The actinopterygian fish Prohalecites from the Triassic of northern Italy: Palaeontology, v. 33, p. 155174.Google Scholar
Tintori, A., 1990b, Dipteronotus olgiatii n. sp. (actynopterigii, Perleidiformes) from the Kalkschieferzone of Cà del Frate (N. Italy): Atti Ticinensi di Scienze della Terra, v. 33, p. 191197.Google Scholar
Tintori, A., 1990c, Estherids from the Kalkschieferzone (Triassic) of Lombardy (N. Italy), in Robba, E., ed., Atti del Quarto Simposio di Ecologia e Paleoecologia delle Comunità Bentoniche: Torino, Museo regionale di Scienze Naturali di Torino, p. 95105.Google Scholar
Tintori, A., 1998, Fish biodiversity in the marine Norian (Late Triassic) of northern Italy: The first Neopterygian radiation: Italian Journal of Zoology, v. 65, p. 193198.CrossRefGoogle Scholar
Tintori, A., 2013, A new species of Saurichthys (Actinopterygii) from the Middle Triassic (early Ladinian) of the northern Grigna Mountain (Lombardy, Italy): Rivista Italiana di Paleontologia e Stratigrafia, v. 119, p. 287302.Google Scholar
Tintori, A., and Brambilla, E., 1991, Sexual dimorphism in a conchostracan population from late Ladinian of southern Calcareous Alps (N. Italy): Contributions from the Palaontological Museum, University of Oslo, v. 364, p. 6566.Google Scholar
Tintori, A., and Lombardo, C., 1999, Late Ladinian fish faunas from Lombardy (N-Italy): Stratigraphy and paleobiology, in Arratia, G., and Schultze, H.P., eds., Mesozoic Fishes 2: Systematics and Fossil Record, München, Verlag Pfeil, p. 495504.Google Scholar
Tintori, A., and Lombardo, C., 2007, A new early Semionotidae (Semionotiformes, Actinopterygii) from the upper Ladinian of Monte San Giorgio area (southern Switzerland and northern Italy): Rivista Italiana di Paleontologia e Stratigrafia, v. 113, p. 369381.Google Scholar
Tintori, A., and Renesto, S., 1983, The Macrosemiidae (Pisces, Actinopterygii) from the Upper Triassic of Lombardy: Rivista Italiana di Paleontologia e Stratigrafia, v. 89, p. 209222.Google Scholar
Tintori, A., and Renesto, S., 1990, A new Lariosaurus from the Kalkschieferzone (Uppermost Ladinian) of Valceresio (Varese, N. Italy): Bollettino della Società Paleontologica Italiana, v. 29, p. 309319.Google Scholar
Tintori, A., Sun, Z.Y., Lombardo, C., Jiang, D.Y., Ji, C., and Motani, R., 2012, A new ‘Flying’ fish from the late Ladinian (Middle Triassic) of Wusha (Guizhou Province, southern China): Gortania, v. 33, p. 3950.Google Scholar
Tintori, A., Jiang, D.Y., Motani, R., Sun, Z.Y., Ji, C., Zou, X.D., and Ma, L.T., 2013, Faunal sequence of Middle Triassic Xingyi Fauna: Evidence from the excavation at Nimaigu of Wusha District, Xingyi City, Guizhou Province: Abstract Book of the 2nd International Symposium on Triassic and Later Marine Vertebrate Faunas, September 10–15, 2013, Xingyi, China, p. 6365.Google Scholar
Tintori, A., Hitij, T., Jiang, D.Y., Lombardo, C., and Sun, Z.Y., 2014, Triassic actinopterygian fishes: The recovery after the end-Permian crisis: Integrative Zoology, v. 9, p. 394411.Google ScholarPubMed
Tintori, A., Lombardo, C., and Kustatscher, E., 2016, The Pelsonian (Anisian, Middle Triassic) fish assemblage from Monte Prà della Vacca/Kühwiesenkopf (Braies Dolomites, Italy): Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, v. 282, p. 181200.CrossRefGoogle Scholar
Van Straelen, V., 1923, Les mysidaces du Callovian de la Boulte-sur- Rhone: Bulletin de la Societé Geologique de France, v. 23, p. 432439.Google Scholar
Wang, X.F., Chen, X.H., Wang, C.S., and Cheng, L., 2009, The Triassic Guanling fossil group: A key GeoPark from a barren mountain, Guizhou Province, China, in Lipps, J.H., and Granier, B.R.C., eds., PaleoParks: The protection and conservation of fossil sites worldwide: Notebooks on Geology, v. 3, p. 1128.Google Scholar
Wirz, A., 1945, Die Triasfauna der Tessiner Kalkalpen. XV Beiträge zur Kenntnis des Ladinikums in Gebiete des Monte San Giorgio: Schweizerische Paläontologische Abhandlungen, v. 65, p. 184.Google Scholar
Zorn, H., 1971, Paläontologische, stratigraphische und sedimentologische Untersuchungen des Salvatoredolomits (Mitteltrias) der Tessiner Kalkalpen: Schweizerische Paläontologische Abhandlungen, v. 91, p. 190.Google Scholar
Zou, X.D., Balini, M., Jiang, D.Y., Tintori, A., Sun, Z.Y., and Sun, Y.L., 2015, The ammonoids from the Zhuganpo Member at Nimaigu and their bearing for dating the Xingyi Fossilägerstatte (Middle Triassic, Guizhou, China): Rivista Italiana di Paleontologia e Stratigrafia, v. 121, p. 135161.Google Scholar