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Bryozoa of the Mission Argillite (Permian), northeastern Washington

  • Ernest H. Gilmour (a1) and Edward M. Snyder (a2)


Fifteen species of Late Permian bryozoans occur in a biohermal bank in the Mission Argillite of northeastern Washington. These include two species conspecific with species described from Japan and 13 new species, one of which is the type species of a new genus. The presence of two species, Dyscritella iwaizakiensis Sakagami, 1961, and Hayasakapora cf. erectoradiata Sakagami, 1960, previously reported from Japan, and the similarity of new species with those previously described from Japan, China and Russia supports the idea that these rocks were originally deposited in the southeastern or central western Pacific Ocean and subsequently accreted to the North American Plate.

Bryozoans and previously reported fusulinids indicate that the biohermal bank is latest Wordian (Kazanian).

Newly described bryozoans include the new genus and type species Sakagamiina easternensis belonging to the Timanodictyidae. Other new species are Fistuliramus pacificus, Meekoporella inflecta, Neoeridotrypella missionensis, Coeloclemis urhausenii, Tabulipora colvillensis, Rhombotrypella kettlensis, Pamirella oculus, Pinegopora petita, Wjatkella nanea, Alternifenestella vagrantia, Polypora arbusca, and Mackinneyella stylettia.



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Astrova, G. G. 1960a. Obshchaya chast, p. 1542. In Fundamentals of Paleontology: Bryozoa and Brachiopoda. Moscow, Akademii Nauk, USSR. (In Russian)
Astrova, G. G. 1960b. Silurijskie Fistuliporidy iz severnykh rajonov RSFSR [Silurian fistuliporids from the northern regions of the RSFSR]. Transactions Akademii Nauk SSSR Kom. Fil. Syktyvkar Institute Geology, 1:352376.
Astrova, G. G. 1964. O novom otryade paleozojskikh mshanok [A new order of Paleozoic Bryozoa]. Paleontologicheskii Zhurnal, 2:2231.
Astrova, G. G. 1965. Morfologiya, istoriya razvitiya i systema ordovikskikhi i silurijskikh mshanok [Morphology, evolutionary history, and systematics of Ordovician and Silurian bryozoans]. Transactions Akademii Nauk SSSR Paleontological Institute, 106:1432.
Astrova, G. G., and Morozova, I. P. 1956. Systematics of Bryozoa of the order Cryptostomata. Doklady Akademii Nauk SSSR, 110:661664. (In Russian)
Bancroft, A. J. 1986. The Carboniferous cystoporate bryozoan Eridopora macrostoma Ulrich from the north of England. Proceedings of the Yorkshire Geological Society, 46:2328.
Bancroft, A. J., and Wyse Jackson, P. N. 1995. Revision of the Carboniferous cystoporate bryozoan Fistulipora incrustans (Phillips, 1836), with remarks on the type species of Fistulipora M'Coy, 1849. Geological Journal, 30:129143.
Bassler, R. S. 1953. Bryozoa, p. G2G253. In Moore, R. C. (ed.), Treatise on Invertebrate Paleontology, Pt. G, Bryozoa. Geological Society of America and University of Kansas Press, Lawrence.
Blake, D. B. 1983. Systematic descriptions for the suborder Rhabdomesina, p. G550G592. In Robison, R. A. (ed.), Treatise on Invertebrate Paleontology, Pt. G, Bryozoa, Revised. Geological Society of America and University of Kansas Press, Lawrence
Boardman, R. S., and Cheetham, A. H. 1983. Glossary of morphological terms, p. G304G320. In Robison, R. A., (ed.), Treatise on Invertebrate Paleontology, Pt. G, Revised. Geological Society of America and University of Kansas Press, Lawrence.
Borg, F. 1926. Studies on Recent cyclostomatous Bryozoa. Zoologiska Bidrag från Uppsala, 10:181507.
Crockford, J. M. 1947. Bryozoa from the Lower Carboniferous of New South Wales and Queensland. Proceedings of the Linnean Society of New South Wales, 72:148.
de Koninck, L. G. 1863. Descriptions of some fossils from India, discovered by Dr. A. Fleming, of Edinburgh. Quarterly Journal of the Geological Survey of London, 19:1.
Dunaeva, N., and Morozova, I. P. 1967. Osobennosti razvitiya i sistematicheskoe polozhenie nekotorykh pozhne-Paleozoyskikh Trepostomat [Evolutionary features and systematic position of some late Paleozoic Trepostomata]. Paleontologicheskii Zhurnal, 4:8694.
Dunbar, C. O. 1953. Parafusulina antimonioensis , p. 1419. In Cooper, G. A., Permian Fauna at El Antimonio, Western Sonora, Mexico. Smithsonian Miscellaneous Collections, v. 119, no. 2.
Ehrenberg, C. G. 1831. Symbolae Physicae, seu Icones et descriptiones Corporum Naturalium novorum aut minus cognitorum, quae ex itineribus per Libyam, Aegyptum, Nubiam, Dongalam, Syriam, Arabiam, et Habessiniam…studio annis 1820-25 redierunt… Pars Zoologica v. 4, Animalia Evertebrata exclusis Insectis. Berolini.
Elias, M. K., and Condra, G. E. 1957. Fenestella from the Permian of West Texas. Geological Society of America Memoir, 70, 158 p.
Gilmour, E. H., McColloch, M. E., and Wardlaw, B. R. 1997. Bryozoa of the Murdock Mountain Formation (Wordian, Permian), Leach Mountains, northeastern Nevada. Journal of Paleontology, 71:214236.
Girty, G. H. 1910. New genera and species of Carboniferous fossils from the Fayetteville Shale of Arkansas. Annals of the New York Academy of Sciences, 20(3):189238.
Gorjunova, R. V. 1975. Permian Bryozoa of the Pamirs. Transactions Paleontological Institute, Akademii Nauk, USSR, 148:1128. (In Russian)
Gorjunova, R. V., and Morozova, I. P. 1979. Late Paleozoic Bryozoa of Mongolia. Trudy Sovmestnaya Sovetski-Mongol'skaya Paleontologicheskaya Ekspeditsiya, 9:1139. (In Russian)
King, W. 1849. On some families and genera of corals. Annals and Magazine of Natural History, 2nd series, 3:388390.
Lee, G. W. 1912. The British Carboniferous Trepostomata. Memoirs of the Geological Survey of Great Britain, 1:135195.
McLaughlin, K. P., and Simons, M. E. 1951. Upper Paleozoic microfossils from Stevens County, Washington. Journal of Paleontology, 25:514519.
M'Coy, F. 1844. A Synopsis of the Characters of the Carboniferous Limestone Fossils of Ireland. Dublin University Press, 207 p.
Miller, M. M. 1987. Dispersed remnants of a northeast Pacific fringing arc: Upper Paleozoic terranes of Permian McCloud faunal affinity, western U. S. Tectonics, 6(6):807830.
Mills, J. W. 1962. High-calcium limestones of eastern Washington. Washington State Division of Mines and Geology Bulletin 48:1268 p.
Mills, J. W., and Davis, J. R. 1962. Permian fossils of the Kettle Falls area, Stevens County, Washington. Contributions from the Cushman Foundation for Foraminiferal Research, 13:4151.
Monger, J. W. H., and Berg, H. C. 1987. Lithotectonic terrane map of western Canada and southeastern Alaska. U.S. Geological Survey Miscellaneous Field Studies Map, MF-1874-B.
Moore, R. C., and Dudley, R. M. 1944. Cheilotrypid bryozoans from Pennsylvanian and Permian rocks of the Midcontinent Region. University of Kansas Publication, State Geological Survey Kansas Bulletin 52:229409.
Morozova, I. P. 1960. The family Eridotrypellidae fam. nov., p. 65. In Fundamentals of Paleontology: Bryozoa and Brachiopoda. Moscow, Akademii Nauk, U.S.S.R. (In Russian)
Morozova, I. P. 1966. A new suborder of Late Palaeozoic bryozoans of the order Cryptostomata. Paleontologicheskii Zhurnal, 2:3341. (In Russian)
Morozova, I. P. 1970. Late Permian Bryozoa. Transactions Paleontological Institute, Akademii Nauk, U.S.S.R., 122:1347. (In Russian)
Morozova, I. P., and Krutchinina, O. N. 1986. Permian Bryozoa of the Arctic, Western Sector. Moscow Science, 144 p.
Morozova, I. P., and Lisitsyn, D. V. 1996. Revision of the Genus Polypora M'Coy. Paleontologicheskii Zhurnal, 4:3847. (In Russian)
Netshcajew, A. V. 1894. The fauna of the Permian deposits of eastern margin of European Russia. Trudy Obshch Estest Voyspit Kazan University 27, 515 p. (In Russian)
Nikiforova, A. I. 1933. Middle Carboniferous Bryozoa of the Donetz Basin. Transactions of the United Geological and Prospecting Service of USSR (Trudy VGRO), Fascile 237, 46 p. (In Russian)
Romantchuk, T. V. 1966. Novye permskie mshanki Khabarorskogo kraya [New Permian bryozoans from Khabarovsk region]. Paleontologicheskii Zhurnal, no. 2:4248.
Romantchuk, T. V. 1970. Hinganella sincera n. sp., p. 9091. In Morozova, I. P. (ed.), Late Permian Bryozoa. Transactions Paleontological Institute, Academii Nauk, U.S.S.R., 122:1347. (In Russian)
Ross, C. A., and Ross, J. R. P. 1983. Late Paleozoic accreted terranes of western North America, p. 722. In Stevens, C. H. (ed.), Pre-Jurassic Rocks in Western North American Suspect Terranes. Los Angeles, California, Pacific Section of Society of Economic Paleontologists and Mineralogists.
Sakagami, S. 1960. Hayasakapora, new Permian bryozoan genus from Iwaizaki, Miyagi Prefecture, Japan. Transactions Proceedings Palaeontological Society Japan, 39:321323.
Sakagami, S. 1961. Japanese Permian Bryozoa. Palaeontological Society Japan, Special Paper, N7:358.
Sakagami, S. 1964. Bryozoa of Akiyoshi, Pt. 1, Permian Bryozoa from the Shigeyasu quarry. Bulletin Akiyoshi-dai Science Museum, Shuhocho, Yamaguchi-ken, Japan, 3:124.
Sakagami, S. 1995. Upper Paleozoic bryozoans from the Lake Titicaca region, Bolivia, Pt. 2, Systematic paleontology. Transactions Proceedings Palaeontological Society of Japan, New Series, 180:261281.
Schulga-Nesterenko, M. I. 1941. Lower Permian Bryozoa of the Urals. Trudy Paleontological Institute, Academy of Science, U.S.S.R., 15, 275 p. (In Russian)
Senter, L. E., Kennedy, D. B., and Gilmour, E. H. 1973. Permian carbonate microfacies of northeastern Washington. Abstracts, Annual Meeting Northwest Scientific Association, p. 7.
Shishova, N. A. 1965. The systematic position and size of the family Hyphamoporidae. Paleontologicheskii Zhurnal, no. 2:5562. (In Russian)
Silberling, N. J., Jones, D. L., Blake, M. C. Jr., and Howell, D. G. 1987. Lithotectonic terrane map of the western conterminous United States. U.S. Geological Survey Miscellaneous Field Studies Map, MF-1874-C.
Simpson, G. B. 1897 (1895). A handbook of the genera of the North American Paleozoic Bryozoa; with an introduction upon the structure of living species. State Geology New York, 14th Annual Report, p. 403669.
Skinner, J. W., and Wilde, G. L. 1966. Permian fusulinids from northeastern Washington, pt. 3 of Permian fusulinids from Pacific Northwest and Alaska. University of Kansas Paleontological Contributions, 4:1622.
Snyder, E. M. 1991. Revised taxonomic procedures and paleoecological applications for some North American Mississippian Fenestellidae and Polyporidae (Bryozoa). Palaeontographica Americana 57, 275 p.
Stevens, C. H. 1995. A giant Permian fusulinid from east-central Alaska with comparisons of all giant fusulinids in western North America. Journal of Paleontology, 69:805812.
Termier, H., and Termier, G. 1971. Bryozoaires du paléozoique supérieur de L'Afghanistan. Documents des Laboratoires de Geology de Faculty Science de Lyon, 47:152.
Ulrich, E. O. 1882. American Paleozoic Bryozoa. Cincinnati Society of Natural History Journal, 5:121175, 232-257.
Utgaard, J. 1983. Paleobiology and taxonomy of the order Cystoporata, p. G327G439. In Robison, R. A., (ed.), Treatise on Invertebrate Paleontology, Pt. G, Bryozoa, Revised. Geological Society of America and University of Kansas Press, Lawrence.
Vine, G. R. 1880. Report of the committee, consisting of Professor P. M. Duncan and Mr. G. R. Vine, appointed for the purpose of reporting on the Carboniferous Polyzoa. British Association for the Advancement of Science, 59th Meeting (Swansea, 1880):7687.
Vine, G. R. 1883. Fourth report of the committee appointed for the purpose of reporting on fossil Polyzoa. Meeting of the British Association for the Advancement of Science, Report 53, 9:161209.
Waagen, W., and Pichl, J. 1885. Salt Range fossils. Palaeontologica Indica, series 13, 1:771814.
Waagen, W., and Wentzel, J. 1886. Salt Range fossils. Palaeontologica Indica, series 13, 1:854892.
Weaver, C. E. 1920. The mineral resources of Stevens County. Washington Geological Survey Bulletin 20:1350.
Xia, Feng-Sheng. 1986. Carboniferous and Early Permian bryozoans from Xainza, northern Xizang. Bulletin of the Nanjing Institute of Geology and Palaeontology. 10:201254.
Yang, Jing-Zhi, and Lu, Lin-Huang. 1984. New material of Early Permian Bryozoa from southwest China. Acta Palaeontologica Sinica, 23:3661.
Young, J. 1883. On Ure's ‘Millepore'; Tabulipora (Cellepora) Urii, Fleming. Annals and Magazine of Natural History Series 5:154158.

Bryozoa of the Mission Argillite (Permian), northeastern Washington

  • Ernest H. Gilmour (a1) and Edward M. Snyder (a2)


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