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Two species of Hesslandona (Phosphatocopida, Crustacea) from the Upper Cambrian of western Hunan, South China and the phylogeny of Phosphatocopida

Published online by Cambridge University Press:  14 July 2015

Huaqiao Zhang ,
Xi-Ping Dong  and
Shuhai Xiao
Huaqiao Zhang
Affiliation:
1School of Earth and Space Sciences, Peking University, Beijing 100871, P. R. China 2State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, P. R. China,
Xi-Ping Dong
Affiliation:
1School of Earth and Space Sciences, Peking University, Beijing 100871, P. R. China 2State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, P. R. China,
Shuhai Xiao
Affiliation:
3Department of Geosciences, Virginia Polytechnic Institute and State University, Blacksburg, VA 24061, USA
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Abstract

The phosphatocopids Hesslandona necopina Müller, 1964 and Hesslandona longispinosa (Kozur, 1974) new combination, recovered from the Upper Cambrian in western Hunan, South China, are described. The ontogenetic stages of H. necopina are revised, with newly defined second and third ontogenetic stages. The second stage is characterized by a bipartite mandibular limb stem consisting of separate coxa and basipod. The third stage is characterized by the partial fusion of the mandibular coxa and basipod. These two parts may become completely fused possibly in a later ontogenetic stage. Vestrogothia longispinosa Kozur, 1974 is reassigned to Hesslandona because of the presence of an interdorsum and a relatively narrow doublure. The new data reported here, plus earlier reports of phosphatocopids from the same section and horizon in western Hunan since 2005, are included in an updated phylogenetic analysis of the Phosphatocopida. Autapomorphies that define several monophyletic groups (e.g., the Crustacea sensu lato, Labrophora, Eucrustacea, Phosphatocopida, Euphosphatocopida, Vestrogothiina, Hesslandonina, and Dorsospinata) are discussed. The present analysis confirms results from earlier phylogenetic analyses in showing the Phosphatocopida-Eucrustacea sister-taxon relationship, but differs from them in supporting the paraphyly of the Hesslandonidae and the monophyly of the Vestrogothiidae.

Type
Research Article
Information
Journal of Paleontology , Volume 85 , Issue 4 , July 2011 , pp. 770 - 788
Copyright
Copyright © The Paleontological Society 

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