Hostname: page-component-76fb5796d-vvkck Total loading time: 0 Render date: 2024-04-26T12:11:27.422Z Has data issue: false hasContentIssue false

Systematics of some Late Ordovician encrinurine trilobites from Laurentian North America

Published online by Cambridge University Press:  14 July 2015

David K. Moss
Affiliation:
Department of Earth Sciences, Syracuse University, Syracuse, NY 13244, USA,
Stephen R. Westrop
Affiliation:
School of Geology and Geophysics, University of Oklahoma, Norman, OK 73072, USA,

Abstract

Classification and relationships of the Ordovician encrinurines Frencrinoides Lespérance and Desbiens and Walencrinoides Lespérance and Desbiens are poorly understood, with little evidence for monophyly of either genus. We revise the type species of both genera, F. capitonis (Frederickson) and W. rarus (Walcott), using new and archival material. We explore their species composition and phylogenetic relationships with a parsimony analysis that includes 17 well-documented ingroup species that can be coded readily, and which is rooted with Encrinuroides regularis Parnaste, the oldest known encrinurine. The results support monophyly of Frencrinuroides and Walencrinuroides, albeit with more limited species membership than proposed by Lespérance and Desbiens. Previous suggestions that both E. uncatus Evitt and Tripp and E. neuter Evitt and Tripp should be assigned to Erratencrinurus Kruger are also supported by our analysis, as is monophyly of Physemataspis Evitt and Tripp. New species are W. rolfi and W. tremblayi.

Type
Research Article
Copyright
Copyright © The Paleontological Society 

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Amati, L. and Westrop, S. R. 2004. A systematic revision of Thaleops (Trilobita: Illaenidae) with new species from the Middle and Late Ordovician of Oklahoma and New York. Journal of Systematic Palaeontology, 2:207256.Google Scholar
Angelin, N. P. 1854. Palaeontologia Suecica, p. 2192. Academiae Regiae Scientarum Suecanae.Google Scholar
Batten Hender, K. L. and Dix, G. R. 2006. Facies, geometry, and geological significance of Late Ordovician (early Caradocian) coral bioherms: Lourdes Formation, western Newfoundland. Sedimentology, 53:13611379.Google Scholar
Batten Hender, K. L. and Dix, G. R. 2008. Facies development of a Late Ordovician mixed carbonate-siliciclastic ramp proximal to the developing Taconic orogen: Lourdes Formation, Newfoundland, Canada. Facies, 54:121149.Google Scholar
Beaumont, C., Quinlan, G., and Hamilton, J. 1988. Orogeny and stratigraphy: Numerical models of the Paleozoic in the eastern interior of North America. Tectonics, 7:389416.Google Scholar
Bergström, S. M., Riva, J., and Kay, M. 1974. Significance of conodonts, graptolites, and shelly faunas from the Ordovician of western and north-central Newfoundland. Canadian Journal of Earth Sciences, 11:16251660.Google Scholar
Bergström, S. M., Chen, X., Gutiérrez-Marco, J. C, and Dronov, A. 2008. The new chronostratigraphic classification of the Ordovician System and its relations to major regional series and stages and to δ13C chemostratigraphy. Lethaia, 41:97107.Google Scholar
Carlucci, J. R. 2012. Trilobite biofacies, systematics and faunal turnover in a sequence stratigraphic framework during the Upper Ordovician of Oklahoma and Virginia. Ph.D. dissertation, University of Oklahoma, Norman, 470 p.Google Scholar
Carlucci, J. R., Westrop, S. R., and Amati, L. 2010. Tetralichine trilobites from the Upper Ordovician of Oklahoma and Virginia, and the phylogenetic systematics of the Tetralichini. Journal of Paleontology, 84:10991120.Google Scholar
Carlucci, J. R., Westrop, S. R., Amati, L., Adrain, J. M., and Swisher, R. E. 2012. A systematic revision of the Upper Ordovician trilobite genus Bumastoides (Illaenidae), with new species from Oklahoma, Virginia and Missouri. Journal of Systematic Palaeontology, 10:679723.Google Scholar
Chatterton, B. D. E. and Ludvigsen, R. 1976. Silicified Middle Ordovician trilobites from the South Nahanni River area, District of Mackenzie, Canada. Palaeontographica Abteilung A, 154, 106 p.Google Scholar
Clarke, J. M. 1894. The lower Silurian trilobites of Minnesota. Geology and Natural History Survey of Minnesota, 3 (2):694759.Google Scholar
Dean, W. T. 1973. Lower Ordovician trilobites from the Summerford Group at Virgin Arm, New World Island, northeastern Newfoundland. Geological Survey of Canada Bulletin 240, 43 p.Google Scholar
Dean, W. T. 1979. Trilobites from the Long Point Group (Ordovician), Port au Port Peninsula, southwestern Newfoundland. Geological Survey of Canada Bulletin 290, 53 p.Google Scholar
Demott, L. L. 1987. Platteville and Decorah trilobites from Illinois and Wisconsin. Minnesota Geological Survey, Report of Investigations, 35:6398.Google Scholar
Desbiens, S. and Lespérance, P. J. 1989. Stratigraphy of the Ordovician of the Lac Saint-Jean and Chicoutimi outliers, Quebec. Canadian Journal of Earth Sciences, 26:11851202.Google Scholar
Edgecombe, G. D. and Chatterton, B. D. E. 1987. Heterochrony in the Silurian radiation of encrinurine trilobites. Lethaia, 20:337351.Google Scholar
Edgecombe, G. D. and Chatterton, B. D. E. 1990. Systematics of Encrinuroides and Curriella (Trilobita), with a new early Silurian encrinurine from the Mackenzie Mountains. Canadian Journal of Earth Sciences, 27:820833.Google Scholar
Edgecombe, G. D., Chatterton, B. D. E., Waisfeld, B. G., and Vaccari, N. E. 1998. Ordovician (Whitrock) calymenid and encrinurid trilobites from the Precordillera of Argentina. Journal of Paleontology, 72:678697.Google Scholar
Evitt, W. R. and Tripp, R. P. 1977. Silicified Middle Ordovician trilobites from the families Encrinuridae and Staurocephalidae. Palaeontographica, Abteilung A, 157:109174.Google Scholar
Fåhraeus, L. E. 1973. Depositional environments and conodont-based correlation of the Long Point Formation (Middle Ordovician), western Newfoundland. Canadian Journal of Earth Sciences, 10:18221833.Google Scholar
Fredrickson, E. A. 1964. Two Ordovician trilobites from southern Oklahoma. Oklahoma Geology Notes, 24:7175.Google Scholar
Goldman, D., S. M. Campbell, and J. M. Rahl. 2002. Three-dimensionally preserved specimens of Amplexograptus (Ordovician, Graptolithina) from the North American midcontinent: taxonomic and biostratigraphic significance. Journal of Paleontology, 76:921927.Google Scholar
Goloboff, P. A., Farris, J. S., and Nixon, K. C. 2008. TNT, a free program for phylogenetic analysis. Cladistics, 24:774786.Google Scholar
Hall, J. 1847. Palaeontology of New York, part VI. Paleontology of New York volume 1: Description of the organic remains from the lower division of the New York System. Carroll and Cook, Albany, New York, 338 p.Google Scholar
Holland, S. M. and Patzkowsky, M. E. 1997. Distal orogenic effects on peripheral bulge sedimentation: Middle and Upper Ordovician of the Nashville Dome. Journal of Sedimentary Research, 67:250263.Google Scholar
Holland, S. M. and Patzkowsky, M. E. 1998. Sequence stratigraphy and relative sea-level of the Nashville Dome, Tennessee. Journal of Sedimentary Research, 68:684699.Google Scholar
Karim, T. and Westrop, S. R. 2002. Taphonomy and paleoecology of Ordovician trilobite clusters, Bromide Formation, south-central Oklahoma. PALAIOS, 17:394402.Google Scholar
Kolata, D. R., Huff, W. D., and Bergström, S. M. 1998. Nature and regional significance of unconformities associated with the Middle Ordovician Hagan K-bentonite complex in the North American midcontinent. Geological Society of America Bulletin, 110:723739.Google Scholar
Krueger, H.-H. 1972. Nachtrag zu “Encrinuriden aus ordovizischen Geschieben” 1971. Geologie, 21:858.Google Scholar
Lespérance, P. J. and Desbiens, S. 1995. Selected Ordovician trilobites from the Lake St. John District of Quebec and their bearing on systematics. Paleontological Society Memoir 42, 19 p.Google Scholar
Ludvigsen, R. 1975. Ordovician formations and faunas, southern Mackenzie Mountains. Canadian Journal of Earth Sciences, 12:663697.Google Scholar
Ludvigsen, R. 1978. Middle Ordovician trilobite biofacies, southern Mackenzie Mountains. Geological Association of Canada Special Paper, 18:137 p.Google Scholar
Ludvigsen, R. 1979. A trilobite zonation of Middle Ordovician rocks, southwestern District of Mackenzie. Geological Survey of Canada Bulletin 312, 99 p.Google Scholar
Mannil, R. 1986. Distribution of trilobites in different facies of the East Baltic Silurian, p. 99109. In Kaljo, D. and Klaaman, E., (eds.), Theory and Practice of Eocostratigraphy. Valgus, Tallinn. (In Russian).Google Scholar
Moss, D. K. 2012. Trilobite faunas and facies of the Upper Ordovician (Sandbian) Lebanon Limestone, Nashville Dome, Tennessee. M.S. thesis, University of Oklahoma, Norman, 177 p.Google Scholar
Nixon, K. C. 2002. WinClada ver. 1.00.08. Published by the author, Ithaca, NY, U.S.A. Google Scholar
Parnaste, H. 2006. The earliest encrinurid trilobites from the East Baltic and their taxonomic interest. Palaeontology, 49:155170.Google Scholar
Phillips, J. and Salter, J. W. 1848. Palaeontological Appendix to Professor John Phillips' Memoir on the Malvern Hills compared with the Paleozoic districts of Abberley, etc. Memoir Geological Survey of Great Britain, 2 (1):331386.Google Scholar
Price, D. 1974. Trilobites from the Sholeshook Limestone (Ashgill) of South Wales. Palaeontology, 17:841868.Google Scholar
Raymond, P. E. and Barton, D. C. 1913. A revision of the American species of Ceraurus . Bulletin of the Museum of Comparative Zoology, 54:525543.Google Scholar
Reed, F. R. C. 1899. The Lower Palaeozoic bedded rocks of County Waterford. The Quarterly Journal of the Geological Society of London, 55:718772.Google Scholar
Reed, F. R. C. 19031906. The Lower Paleozoic trilobites of the Girvan area Ayrshire. Palaeontographical Society Monograph, 186 p.Google Scholar
Reed, F. R. C. 1914. The Lower Paleozoic trilobites of the Girvan area Ayrshire. Palaeontographical Society Monograph, Supplement, 73 p.Google Scholar
Reed, F. R. C. 1931. The Lower Paleozoic trilobites of the Girvan area Ayrshire. Palaeontographical Society Monograph, Supplement 2: 30 p.Google Scholar
Safford, J. M. 1869. Geology of Tennessee. Nashville, 550 p.Google Scholar
Safford, J. M. and Vodges, A. W. 1889. Description of new species of fossil Crustacea from the lower Silurian of Tennessee, with remarks on other not well known. Proceedings of the Academy of Natural Sciences of Philadelphia, 41:166168.Google Scholar
Shaw, F. C. 1968. Early Middle Ordovician Chazy trilobites of New York. New York State Museum and Science Service, Memoir 17, 163 p.Google Scholar
Shaw, F. C. 1974. Simpson Group (Middle Ordovician) trilobites of Oklahoma. Paleontological Society Memoir, 6, 54 p.Google Scholar
Stait, B. and Barnes, C. R. 1991. Stratigraphy of the Middle Ordovician Long Point Group, western Newfoundland. Geological Survey of Canada Paper 90-9:235244.Google Scholar
Strong, E. E. and Lipscomb, D. 1999. Character coding and inapplicable characters. Cladistics, 15:363371.Google Scholar
Strusz, D. L. 1980. The Encrinuridae and related trilobite families, with a description of Silurian species from southeastern Australia. Palaeontographica Abteilung A, 168, 68 p.Google Scholar
Sutherland, P. K. and Amsden, T. W., 1959. A re-illustration of the trilobite Lonchodomas mcgeheei Decker from the Bromide Formation (Orovician) of southern Oklahoma. Oklahoma Geology Notes, 19:212219.Google Scholar
Swofford, D. L. 2001. PAUP∗. Phylogenetic Analysis Using Parsimony Version 4. Sinauer Associates, Sunderland, Massachusetts.Google Scholar
Templeton, J. S. and Willman, H. B., 1963. Champlainian Series (Middle Ordovician) in Illinois. Illinois Geological Survey Bulletin 89, 259 p.Google Scholar
Tremblay, J. V. and Westrop, S. R. 1991. Middle Ordovician (Whiterockian) trilobites from the Sunblood Formation, District of Mackenzie, Canada. Journal of Paleontology, 65:801824.Google Scholar
Troedsson, G. F. 1928. On the Middle and Upper Ordovician faunas of Northern Greenland Part II. Meddelelser om Gr⊘nland, 72:1198.Google Scholar
Tripp, R. P. 1962. Trilobites from the confinis flags (Ordovician) of the Girvan district, Ayrshire. Transactions of the Royal Society of Edinburgh, 65:136.Google Scholar
Tripp, R. P. 1965. Trilobites from the Albany division (Ordovician) of the Girvan district, Ayrshire. Palaeontology, 8:577603.Google Scholar
Tripp, R. P. 1967. Trilobites form the Upper Stinchar Limestone (Ordovician) of the Girvan district, Ayrshire. Transactions of the Royal Society of Edinburgh, 67:4393.Google Scholar
Tripp, R. P. 1979. Trilobites form the Ordovician Auchensoul and Stinchar limestones of the Girvan District, Strathclyde. Palaeontology, 22:339361.Google Scholar
Tripp, R. P. 1980. Trilobites from the Ordovician Ardwell Group of the Craighead Inlier, Girvan District, Scotland. Transactions of the Royal Society of Edinburgh: Earth Sciences, 71:147157.Google Scholar
Walcott, C. D. 1877 [1897]. Descriptions of new species of fossil from the Chazy and Trenton limestones. Annual Report on the New York State Museum of Natural History by the Regents of the University of the State of New York, 31:6871.Google Scholar
Westrop, S. R. and Adrain, J. M. 2007. Bartonaspis new genus, a trilobite species complex from the base of the upper Cambrian Sunwaptan Stage in North America. Canadian Journal of Earth Sciences, 44:9871003.Google Scholar
Whittington, H. B. 1950. Sixteen Ordovician genotype trilobites. Journal of Paleontology, 24:531565.Google Scholar
Whittington, H. B. 1965. The Ordovician trilobites of the Bala area, Merioneth. Palaeontographical Society Monograph, p. 93138.Google Scholar