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New mud turtles (Kinosternidae, Kinosternon) from the middle–late Miocene of the United States

Published online by Cambridge University Press:  21 March 2016

Jason R. Bourque
Division of Vertebrate Paleontology, Florida Museum of Natural History, University of Florida, Gainesville, Florida 32611, USA 〈〉
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Kinosternon Spix, 1824, was widespread in xeric to savanna associated paleowetlands across North America during the middle–late Miocene and steadily diversified following its first occurrences in the Hemingfordian. In the middle Miocene, Kinosternon rincon n. sp. occurred in the late Barstovian Cerro Conejo Formation of north central New Mexico, perhaps concurrently with Kinosternon pojoaque Bourque 2012a from the Tesuque Formation. Subsequent late Miocene kinosternine fossils indicate that at least three potentially contemporaneous species existed throughout the Clarendonian. These are Kinosternon pannekollops n. sp., from the Ogallala Formation of northern Texas; Kinosternon wakeeniense, n. sp., from the Ogallala Formation of northwestern Kansas and Ash Hollow Formation of south-central Nebraska; and Kinosternon notolophus n. sp., from the Alachua and Statenville formations of northern Florida. Kinosternon rincon is phylogenetically nested between the Kinosternon flavescens (Agassiz, 1857) group (=Platythyra Agassiz, 1857) and more derived Kinosternon including the Kinosternon subrubrum (Lacépède, 1788) group (=Thyrosternum Agassiz, 1857). Kinosternon pannekollops is recovered on the stem of the K. subrubrum group and is the oldest and largest member of that lineage. Kinosternon notolophus is readily differentiated from other Miocene Kinosternon in possessing a distinct dorsomedial keel on the nuchal and faint dorsolateral costal carination. The K. subrubrum group probably originated in the late Miocene savannas of the Great Plains and dispersed eastward via the Gulf Coastal Plain. An unnamed kinosternine taxon existed in the coastal plains of the eastern and southeastern United States during the Middle Miocene Climatic Optimum (with fossils from ~18 to 15 Ma), and disappeared from the region coincident with the end of that megathermal event.

Copyright © 2016, The Paleontological Society 

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Adler, K., 1968, Synonymy of the Pliocene turtles Pseudemys hilli Cope and Chrysemys limnodytes Galbreath: Journal of Herpetology, v. 1, p. 3238.CrossRefGoogle Scholar
Agassiz, L., 1857, Contributions to the Natural History of the United States of America, First Monograph. Volume 1, Part 2, North American Testudinata. Volume 2, Part 3, Embryology of the Turtle: Boston, Little, Brown and Company, p. 233643.Google Scholar
Berry, J.F., and Iverson, J.B., 1980, A new species of mud turtle, genus Kinosternon, from Oaxaca, Mexico: Journal of Herpetology, v. 14, p. 313320.Google Scholar
Berry, J.F., and Legler, J.M., 1980, A new turtle (genus Kinosternon) from Sonora, Mexico: Contributions in Science, Los Angeles County Museum, v. 352, p. 112.Google Scholar
Berry, J.F., Seidel, M.E., and Iverson, J.B., 1997, A new species of mud turtle (genus Kinosternon) from Jalisco and Colima, Mexico, with notes on its natural history: Chelonian Conservation and Biology, v. 2, p. 329337.Google Scholar
Bourque, J.R., 2011, Reassessment of a putative fossil stinkpot (Kinosternidae: Sternotherus) from the late Miocene (Clarendonian) of Kansas: Journal of Herpetology, v. 45, p. 234237.CrossRefGoogle Scholar
Bourque, J.R., 2012a, An extinct mud turtle of the Kinosternon flavescens group (Testudines, Kinosternidae) from the middle Miocene (late Barstovian) of New Mexico: Journal of Vertebrate Paleontology, v. 32, p. 6881.CrossRefGoogle Scholar
Bourque, J.R., 2012b, A fossil mud turtle (Testudines, Kinosternidae) from the early middle Miocene (early Barstovian) of New Mexico: Journal of Vertebrate Paleontology, v. vol. 32, p. 836853.CrossRefGoogle Scholar
Bourque, J.R., 2013, Fossil Kinosternidae from the Oligocene and Miocene of Florida, USA, in Brinkman, D.B., Holroyd, P.A., and Gardner, J.D., eds., Morphology and Evolution of Turtles. Vertebrate Paleobiology and Paleoanthropology Series: Dordrecht, Netherlands, Springer, p. 459475.Google Scholar
Bourque, J.R., and Schubert, B.W., 2015, Fossil musk turtles (Kinosternidae, Sternotherus) from the late Miocene–early Pliocene (Hemphillian) of Tennessee and Florida: Journal of Vertebrate Paleontology, v. 35, doi:10.1080/02724634.2014.885441.CrossRefGoogle Scholar
Bramble, D.M., Hutchison, J.H., and Legler, J.M., 1984, Kinosternid shell kinesis: Structure, function, and evolution: Copeia, v. 1984, p. 456475.CrossRefGoogle Scholar
Bryant, J.D., 1991, New early Barstovian (middle Miocene) vertebrates from the upper Torreya Formation, eastern Florida panhandle: Journal of Vertebrate Paleontology, v. 11, p. 472489.CrossRefGoogle Scholar
Connell, S.D., 2001, Stratigraphy of the Albuquerque Basin, Rio Grande Rift, central New Mexico: A progress report, in Connell, S.D., Lucas, S.G., and Love, D.W., eds., Stratigraphy and Tectonic Development of the Albuquerque Basin, Central Rio Grande Rift. Field-Trip Guidebook for the Geological Society of America, Mini-Papers: Socorro, New Mexico, New Mexico Bureau of Mines and Mineral Resources, p. A1A27.Google Scholar
Connell, S.D., Koning, D.J., Kelley, S.A., and Brandes, N.N., 2007, Oligocene and early Miocene sedimentation in the southwestern Jemez Mountains and northwestern Albuquerque Basin, New Mexico, in Kues, B.S., Kelley, S.A., and Lueth, V.W., eds., New Mexico Geological Society Guidebook, 58th Field Conference, Geology of the Jemez Mountains Region II: Socorro, New Mexico, New Mexico Bureau of Mines and Mineral Resources, p. 195–208.Google Scholar
Cope, E.D., 1878, Descriptions of new extinct Vertebrata from the upper Tertiary and Dakota formations: Bulletin of the United States Geological and Geographical Survey, Territories, v. 4, p. 379396.Google Scholar
Duméril, A.M.C., and Duméril, A.H.A., 1851, Catalogue methodique de la Collection des Reptiles du Museum d’Histoire Naturelle: Paris, Gide and Boudry, p. iv + 224.Google Scholar
Ehrenfeld, J.G., and Ehrenfeld, D.W., 1973, Externally secreting glands of freshwater and sea turtles: Copeia, v. 1973, p. 305314.CrossRefGoogle Scholar
Ernst, C.H., and Lovich, J.E., 2009, Turtles of the United States and Canada, 2d Ed. Baltimore, Maryland, The Johns Hopkins University Press, 827 p.Google Scholar
Eronen, J.T., Fortelius, M., Micheels, A., Portmann, F.T., Puolamäki, K., and Janis, C.M., 2013, Neogene aridification of the Northern Hemisphere: Geology, v. 40, p. 823836.CrossRefGoogle Scholar
Fichter, L.S., 1969, Geographical distribution and osteological variation in fossil and Recent specimens of two species of Kinosternon (Testudines): Journal of Herpetology, v. 3, p. 113119.CrossRefGoogle Scholar
Fitzinger, L., 1835, Entwurf einer systematischen Anordnung der Schildkröten nach den Grundsätzen der natürlichen Methode: Annalen des Naturhistorischen Museums in Wien, v. 1, p. 103128.Google Scholar
Galbreath, E.C., 1948, A new extinct emydid turtle from the Lower Pliocene of Oklahoma: University of Kansas Publications, Museum of Natural History, v. 1, p. 269275.Google Scholar
Garman, S., 1891, On a tortoise found in Florida and Cuba, Cinosternum baurii: Bulletin of the Essex Institute, v. 23, p. 141144.Google Scholar
Geoffroy Saint-Hilaire, E., 1809, Sur les tortues molles, nouveau genre sous le nom Trionyx, et sur la formation des carapaces: Annales Muséum National d’Histoire Naturelle, v. 14, p. 120.Google Scholar
Gidley, J.W., 1903, A new three-toed horse: Bulletin of the American Museum of Natural History, v. 19, p. 465476.Google Scholar
Gilmore, C.W., 1922, A new fossil turtle, Kinosternon arizonense, from Arizona: Proceedings of the United States National Museum, v. 62, p. 18.CrossRefGoogle Scholar
Glass, B., and Hartweg, N., 1951, Kinosternon murrayi, a new musk turtle of the hirtipes group from Texas: Copeia, v. 1951, p. 5052.CrossRefGoogle Scholar
Gray, J.E., 1825, A synopsis of the genera of reptiles and amphibia, with a description of some new species: Annals of Philosophy, v. 10, p. 193217.Google Scholar
Gray, J.E., 1831, Synopsis Reptilium or Short Descriptions of the Species of Reptiles. Part I. Cataphracta, Tortoises, Crocodiles, and Enaliosaurians: London, Treuttel, Wurz Company, p. viii + 85.CrossRefGoogle Scholar
Gray, J.E., 1844, Catalogue of the Tortoises, Crocodiles, and Amphisbaenians in the Collection of the British Museum: London, 80 p.Google Scholar
Gray, J.E., 1856a, On some new species of freshwater tortoises from North America, Ceylon, and Australia, in the collection of the British Museum: Proceedings of the Zoological Society London, v. 1855, p. 197202.Google Scholar
Gray, J.E., 1856b, Catalogue of the Shield Reptiles in the Collection of the British Museum, Part 1: Testudinata (Tortoises): London, Taylor and Francis, 198 p.Google Scholar
Hartweg, N., 1934, Description of a new kinosternid from Yucatan: Occasional Papers of the Museum of Zoology, University of Michigan, v. 277, p. 12.Google Scholar
Hay, O.P., 1892, The batrachians and reptiles of the State of Indiana: Annual Report of the Indiana Department of Geology and Natural Resources, v. 17, p. 409602.Google Scholar
Hay, O.P., 1906, Descriptions of two new genera (Echmatemys and Xenochelys) and two new species (Xenochelys formosa and Terrapene putnami) of fossil turtles: Bulletin of the American Museum of Natural History, v. 22, p. 2731.Google Scholar
Hay, O.P., 1908, The fossil turtles of North America: Carnegie Institution of Washington, Publication no. 75, 568 p., +113 pl.Google Scholar
Holman, J.A., 1975, Herpetofauna of the WaKeeney local fauna (lower Pliocene: Clarendonian) of Trego County, Kansas: University of Michigan Papers on Paleontology, v. 12, p. 4966.Google Scholar
Holman, J.A., 1981, Fossil amphibians and reptiles of Nebraska and Kansas: National Geographic Society Research Reports, v. 13, p. 253262.Google Scholar
Holman, J.A., 1998, Reptiles of the lower Miocene (Hemingfordian) Pollack Farm Fossil Site, Delaware: in Benson, R.N., ed., Geology and Paleontology of the Lower Miocene Pollack Farm Fossil Site: Delaware, Delaware Geological Survey, Special Publication 21, University of Delaware, p. 141147.Google Scholar
Hulbert, R.C. Jr., 2001, The Fossil Vertebrates of Florida: Gainesville, University Press of Florida, 351 p.Google Scholar
Hutchison, J.H., 1991, Early Kinosterninae (Reptilia: Testudines) and their phylogenetic significance: Journal of Vertebrate Paleontology, v. 11, p. 145167.CrossRefGoogle Scholar
Hutchison, J. H., and Bramble, D. M., 1981, Homology of the plastral scales of the Kinosternidae and related turtles: Herpetologica, v. 37, p. 7385.Google Scholar
Iverson, J.B., 1979, A taxonomic reappraisal of the yellow mud turtle, Kinosternon flavescens (Testudines: Kinosternidae): Copeia, v. 1979, p. 212225.CrossRefGoogle Scholar
Iverson, J. B., 1991, Phylogenetic hypotheses for the evolution of modern kinosternine turtles: Herpetological Monographs, v. 5, p. 127.CrossRefGoogle Scholar
Iverson, J.B., 1998, Molecules, morphology, and mud turtle phylogenetics (family Kinosternidae): Chelonian Conservation and Biology, v. 3, p. 113117.Google Scholar
Iverson, J.B., Brown, R.M., Akre, T.S., Near, T.J., Le, M., Thomson, R.C., and Starkey, D.E., 2007, In search of the tree of life for turtles, in Schaffer, H.B., FitzSimmons, N.F., Georges, A., and Rhodin, A.G.H., eds., Defining Turtle Diversity: Proceedings of a Workshop on Genetics: Ethics, and Taxonomy of Freshwater Turtles and Tortoises, Chelonian Research Monographs, v. 4, p. 85106.Google Scholar
Iverson, J.B., Le, M., and Ingram, C., 2013, Molecular phylogenetics of the mud and musk turtle family Kinosternidae: Molecular Phylogenetics and Evolution, v. 69, p. 929939.CrossRefGoogle ScholarPubMed
Jackson, D.R., 1978, Evolution and fossil record of the chicken turtle Deirochelys, with a re-evaluation of the genus: Tulane Studies in Zoology and Botany, v. 20, p. 3555.Google Scholar
Lacépède, B., 1788, Histoire Naturelle des Quadrupèdes Ovipares et des Serpens Vol. 1: Paris, Hôtel de Thou, p. 651.CrossRefGoogle Scholar
Latreille, P.A., 1801, Histoire Naturelle des Reptiles, in Sonnini, C.S., and Latreille, P.A., eds., Histoire Naturelle des Reptiles, avec figures dessinées d’après nature. Tome Premier. Première Partie. Quadrupèdes et Bipèdes Ovipares: Paris, Imprimerie de Crapelet, p. 280.Google Scholar
LeConte, J., 1854, Description of four new species of Kinosternum: Proceedings of the Academy of Natural Sciences of Philadelphia, v. 7, p. 180190.Google Scholar
Legler, J.M., 1965, A new species of turtle, genus Kinosternon, from Central America: Lawrence, University of Kansas Publications, Museum of Natural History, v. 15, p. 615625.CrossRefGoogle Scholar
Leidy., J., 1858, Notice of remains of extinct Vertebrata, from the valley of the Niobrara River, collected during the exploring expedition of 1857, in Nebraska, under the command of Lieutenant G. K. Warren, by Dr. F. V. Hayden, geologist to the expedition: Proceedings of the Academy of Natural Sciences Philadelphia, v. 10, p. 20–29.Google Scholar
Linnaeus, C., 1758, Systema Naturae, per Regna Tria Naturae, secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. Tomus I. Editio Decima, Reformata: Stockholm, Lorentii Salvii, p. 824.Google Scholar
Linnaeus, C., 1766, Systema Naturae. Editio Duodecima, Reformata. Tomus I, Pars I, Regnum Animale: Stockholm, Laurentii Salvii, 532 p.Google Scholar
Maddison, W.P., and Maddison, D.R., 2008, Mesquite: a modular system of evolutionary analysis, version 2.5: http// (accessed February 2016).Google Scholar
Merrem, B., 1820, Tentamen Systematis Amphibiorum: Marburg, Germany: Krieger, p. 188.Google Scholar
Morgan, G.S., 1989, Miocene vertebrate faunas from the Suwannee River Basin of north Florida and south Georgia, in Morgan, G.S., ed., Miocene Paleontology and Stratigraphy of the Suwannee River Basin of North Florida and South Georgia: Tallahassee, Florida, Southeastern Geological Society, Guidebook, v. 30, p. 2653.Google Scholar
Morgan, G.S., and Lucas, S.G., 2001, Miocene mammalian faunas and biostratigraphy of the Zia Formation, northern Albuquerque Basin, Sandoval County, New Mexico, in Connell, S.D., Lucas, S.G., and Love, D.W., eds., Stratigraphy and Tectonic Development of the Albuquerque Basin, Central Rio Grande Rift. Field-Trip Guidebook for the Geological Society of America, Mini-Papers: Socorro, New Mexico, New Mexico Bureau of Mines and Mineral Resources, p. C33C35.Google Scholar
Neill, W.T., 1948, Use of scent glands by prenatal Sternotherus minor: Herpetologica, v. 4, p. 148.Google Scholar
Parmley, D., 1992, Turtles from the late Hemphillian (latest Miocene) of Knox County, Nebraska: Texas Journal of Science, v. 44, p. 339348.Google Scholar
Plummer, M.V., and Trauth, S.E., 2009, The structure of Rathke’s glands in the softshell turtles Apalone mutica and A. spinifera: Herpetological Conservation and Biology, v. 4, p. 207220.Google Scholar
Pound, M.J., Haywood, A.M., Salzmann, U., Riding, J.B., Lunt, D.J., and Hunter, S.J., 2011, A Tortonian (Late Miocene, 11.61–7.25) global vegetation reconstruction: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 300, p. 2945.CrossRefGoogle Scholar
Pound, M.J., Haywood, A.M., Salzmann, U., and Riding, J.B., 2012, Global vegetation dynamics and latitudinal temperature gradients during the mid to late Miocene: Earth-Science Reviews, v. 112, p. 122.CrossRefGoogle Scholar
Rafinesque, W.J., 1832, Descriptions of two new genera of soft-shell turtles of North America: Atlantic Journal and Friend of Knowledge, v. 1, p. 6465.Google Scholar
Reid, B.N., Le, M., McCord, W.P., Iverson, J.B., Georges, A., Bergmann, T., Amato, G., DeSalle, R., and Naro-Maciel, E., 2011, Comparing and combining distance-based and character-based approaches for barcoding turtles: Molecular Ecology Resources, v. 11, p. 956967.CrossRefGoogle ScholarPubMed
Rose, F.L., and Weaver, W.G. Jr., 1966, Two new species of Chrysemys (=Pseudemys) from the Florida Pliocene: Tulane Studies in Geology, v. 5, p. 4148.Google Scholar
Schmidt, K.P., 1947, A new kinosternid turtle from Colombia: Fieldiana: Zoology, v. 31, p. 109112.Google Scholar
Schultz, G.E., 1977, Guidebook: Field Conference on Late Cenozoic Biostratigraphy of the Texas Panhandle and Adjacent Oklahoma, August 4–6, 1977. Killgore Research Center, Department of Geology and Anthropology, Special Publication No. 1: Canyon, Texas, West Texas State University, 160 p.Google Scholar
Schultz, G.E., 1990, Clarendonian and Hemphillian vertebrate faunas from the Ogallala Formation (late Miocene–early Pliocene) of the Texas Panhandle and adjacent Oklahoma, in Gustavson, T.C., ed., Geologic Framework and Regional Hydrology: Upper Cenozoic Blackwater Draw and Ogallala Formations, Great Plains Austin: Bureau of Economic Geology, University of Texas, p. 5697.Google Scholar
Schultz, G.E., 2002, Clarendonian and Hemphillian vertebrate faunas from the Ogallala Formation (late Miocene–early Pliocene) of the Texas Panhandle and adjacent Oklahoma, in Burkhalter, R., Czaplewski, N., and Lupia, R., eds., Field Trip Guidebook for the 62nd Annual Meeting of the Society of Vertebrate Paleontology, Norman, Oklahoma. Oklahoma Geological Survey Open-File Report 10-2002: Norman, University of Oklahoma, p. 3371.Google Scholar
Schweigger, A.F., 1812, Monographiae Cheloniorum: Königsberger Archiv für Naturwissenschaften und Mathematik, v. 1, p. 271368. 406–458.Google Scholar
Seidel, M.E., Iverson, J.B., and Adkins, M.D., 1986, Biochemical comparisons and phylogenetic relationships in the family Kinosternidae (Testudines): Copeia, v. 1986, p. 285294.CrossRefGoogle Scholar
Siebenrock, F., 1906, Eine neue Cinosternum-Art aus Florida: Zoologischer Anzeiger, v. 30, p. 727728.Google Scholar
Spinks, P.Q., Thomson, R.C., Gidis, M., and Shaffer, H.B., 2014, Multilocus phylogeny of the New-World mud turtles (Kinosternidae) supports the traditional classification of the group: Molecular Phylogenetics and Evolution, v. 76, p. 254260.CrossRefGoogle ScholarPubMed
Spix, J.B. von, 1824, Animalia Nova, Species Novae Testudinum et Ranarum quas in Itinere per Brasiliam Annis 1817–1820 Jussu et Auspicis Maximiliani Josophi I. Bavariae Regis: Munich, Hübschmann, v. 53, p.Google Scholar
Stejneger, L., 1925, New species and subspecies of American turtles: Journal of the Washington Academy of Sciences, v. 15, p. 462463.Google Scholar
Swofford, D.L., 2003, PAUP* 4.0-b10: Sunderland, Massachusetts, Sinauer Associates.Google Scholar
Tedford, R.H., 1981, Mammalian biochronology of the late Cenozoic basins of New Mexico: Geological Society of America Bulletin, v. 92, p. 10081022.2.0.CO;2>CrossRefGoogle Scholar
Tedford, R.H., Albright, L.B. III, Barnosky, A.D., Ferrusquia-Villafranca, I., Hunt, R.M. Jr., Storer, J.E., Swisher, C.C. III, Voorhies, M.R., Webb, S.D., and Whistler, D.P., 2004, Mammalian biochronology of the Arikareean through Hemphillian interval (late Oligocene through early Pliocene epochs), in Woodburne, M.O., ed., Late Cretaceous and Cenozoic Mammals of North America: Biostratigraphy and Geochronology: New York, Columbia University Press, p. 169231.Google Scholar
Wagler, J.G., 1830, Natürliches System der Amphibien, mit vorangehender Classification der Säugethiere und Vögel. Ein Beitrag zur vergleichenden Zoologie: Munich, J.G. Cotta’schen, Buchhandlung, 354 p.Google Scholar
Webb, S.D., 1977, A history of savanna vertebrates in the New World. Part 1: North America: Annual Review of Ecology and Systematics, v. 8, p. 355380.CrossRefGoogle Scholar
Webb, S.D., MacFadden, B.J., and Baskin, J.A., 1981, Geology and paleontology of the Love Bone Bed from the late Miocene of Florida: American Journal of Science, v. 281, p. 513544.CrossRefGoogle Scholar
Weems, R.E., and George, R.A., 2013, Amphibians and nonmarine turtles from the Miocene Calvert Formation of Delaware, Maryland, and Virginia (USA): Journal of Paleontology, v. 87, p. 570588.CrossRefGoogle Scholar
Wilson, R.L., 1968, Systematics and faunal analysis of a lower Pliocene vertebrate assemblage from Trego County, Kansas: Contributions from the Museum of Paleontology, University of Michigan, v. 22, p. 75126.Google Scholar
Zachos, J., Pagani, M., Sloan, L., Thomas, E., and Billups, K., 2001, Trends, rhythms, and aberrations in global climate 65 Ma to present: Science, v. 292, p. 686693.CrossRefGoogle ScholarPubMed
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