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Cambrian microfossils from the Tethyan Himalaya

Published online by Cambridge University Press:  15 June 2016

Ian R. Gilbert
Department of Earth Sciences, University of California, Riverside, CA 92521, USA 〈〉, 〈〉
Nigel C. Hughes
Department of Earth Sciences, University of California, Riverside, CA 92521, USA 〈〉, 〈〉
Paul M. Myrow
Department of Geology, Colorado College, Colorado Springs, CO 80903, USA 〈〉


Cambrian biostratigraphy of the Indian subcontinent is best documented from the Parahio Formation of the Tethyan Himalaya. Recently established trilobite biostratigraphy shows that the formation encompasses the latest part of unnamed Stage 4 and much of unnamed Stage 5. A variety of small shelly fossils have been recovered via acid digestion of carbonate beds and include tetract and pentact hexactinellid sponge spicules, chancelloriid spicules belonging to Chancelloria sp. and a new species, Archiasterella dhiraji, shells of an helcionelloid comparable to Igorella maidipingensis, a meraspid ptychopariid trilobite, the tubular Cupitheca sp., a poorly preserved hyolith, and an assortment of spinose microfossils of uncertain affinity. These newly recovered microfossils are consistent with the trilobite-based lower and middle Cambrian age determination and do not support a late Cambrian age for the top of the Parahio Formation advocated in some recent literature. The microfossils reported herein significantly expand the known diversity of such fossils from Cambrian strata in the Himalayan region, and allow for comparison of this fauna with others from Gondwanaland and elsewhere. Integration with trilobite data indicate that the stratigraphic ranges of many small shelly fossils described in this study are greater than previously recognized.

Copyright © 2016, The Paleontological Society 

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Abrantes, F., Gil, I., Lopes, C., and Castro, M., 2005, Quantitative diatom analyses: a faster cleaning procedure: Deep Sea Research Part I: Oceanographic Research Papers, v. 52(1), p. 189198.CrossRefGoogle Scholar
Bengtson, S., and Missarzhevsky, V.V., 1981, Coeloscleritophora: a major group of enigmatic Cambrian metazoans, in Taylor, M.E., ed.., Short papers from the Second International Symposium on the Cambrian System: U.S. Geological Survey Open-File Report, 81-743, p. 1921.Google Scholar
Bengtson, S., Conway Morris, S., Cooper, B.J., Jell, P.A., and Runnegar, B.N., 1990, Early Cambrian fossils from South Australia: Memoirs of the Association of Australian Palaeontologists, v. 9, p. 1364.Google Scholar
Bhatt, D.K., and Kumar, G., 1980, Discovery of conodonts in the Cambrian of Spiti, Tethys Himalaya: Current Science, v. 49, p. 357358.Google Scholar
Bhatt, D.K., Mamgain, V.D., and Misra, R.S., 1985, Small shelly fossils of Early Cambrian (Tommotian) age from Chert-Phosphorite Member, Tal Formation, Mussoorie syncline, Lesser Himalaya, India and their chronostratigraphic evaluation: Journal of the Palaeontological Society of India, v. 30, p. 92102.Google Scholar
Brasier, M.D., 1989, Towards a biostratigraphy of the earliest skeletal biotas, in Cowie, J.W., and Brasier, M.D., eds., The Precambrian–Cambrian boundary, Oxford, Clarendon Press, p. 117165.Google Scholar
Brasier, M.D., and Singh, P., 1987, Microfossils and Precambrian–Cambrian boundary stratigraphy at Maldeota, Lesser Himalaya: Geological Magazine, v. 124, p. 323345.CrossRefGoogle Scholar
Cederström, P., Ahlberg, P., Nilsson, C.H., Ahlgren, J., and Eriksson, M.E., 2011, Moulting, ontogeny and sexual dimorphism in the Cambrian ptychopariid trilobites Strenuaeva inflata from the northern Swedish Caledonides: Palaeontology, v. 54, p. 685703.CrossRefGoogle Scholar
Chen, Y., and Zhang, S., 1980, Small shelly fossils from the Early Lower Cambrian, Sonlingpo, Eastern Yangtze Gorges: Geological Review, v. 26(3), p. 190197. [in Chinese]Google Scholar
Cuvier, G., 1797, Tableau Élementaire de l’historie Naturelle des Animaux, Paris, 710 p.Google Scholar
Devaere, L., Clausen, S., Steiner, M., Álvaro, J.J., and Vachard, D., 2013, Chronostratigraphic and palaeogeographic significance of an early Cambrian microfauna from the Heraultia Limestone, northern Montagne Noire, France: Palaeontologia Electronica, v. 16.2, p. 191.Google Scholar
Dong, X., and Knoll, A.H., 1996, Middle and late Cambrian sponge spicules from Hunan, China: Journal of Paleontology, v. 70(2), p. 173184.Google Scholar
Doré, F., and Reid, R.E., 1965, Allonnia tripodophora nov. gen., nov. sp., nouvelle ponge du Cambrien Infrieur de Carteret (Manche): Compte Rendu Sommaire des Séances de a Société Géologique de France, v. 1965, p. 2021.Google Scholar
Duan, C., 1984, Small shelly fossils from the Lower Cambrian Xihaoping Formation in the Shennongjia District, Hubei Province—hyoliths and fossil skeletons of unknown affinities: Bulletin of the Tianjin Institute of Geology and Mineral Resources, v. 7, p. 143188.Google Scholar
Esakova, N.V., and Zhegallo, E.A., 1996, Biostratigraphy and fauna of the Lower Cambrian of Mongolia: Trudy, Sovmestnaya Rossiysko-Mongol’skaya Paleontologicheskaya Ekspeditsiya, v. 46, p. 1214.Google Scholar
Fernández Remolar, D.C., 2001, Chancelloridae del Ovetiense Inferior de la Sierra de Córdoba, España: Revista Española de Paleontología, v. 16(1), p. 3961.Google Scholar
Fuchs, G.R., and Mostler, H., 1972, Der erste Nachweis von Fossilien (Kambrischen Alters) in der Hazira Formation, Hazara, Pakistan: Geologisch-Paläontologische Mitteilungen, Innsbruck, v. 2, p. 112.Google Scholar
Gehling, J.G., and Rigby, J.K., 1996, Long expected sponges from the Neoproterozoic Ediacara fauna of South Australia: Journal of Paleontology, v. 70.2, p. 185195.CrossRefGoogle Scholar
Geyer, G., 1994, Middle Cambrian molluscs from Idaho and early conchiferan evolution: New York State Museum Bulletin, v. 481, p. 6986.Google Scholar
Grant, R.E., 1836, Animal kingdom: The Cyclopaedia of Anatomy and Physiology, v. 1, p. 107118.Google Scholar
Gravestock, D.I., Alexander, E.M., Demidenko, Y.E., Esakova, N.V., Holmer, L.E., Jago, J.B., Lin, T.-R., Mel’nikova, L.M., Parkhaev, P.Yu., Rozanov, P.Yu., Ushatinskaya, G.T., Zang, Y.-L., Zhegallo, E.A., and Zhuravlev, A.Yu., 2001, The Cambrian Biostratigraphy of the Stansbury Basin, South Australia: Transactions of the Palaeontological Institute, Russian Academy of Sciences, v. 282, p. 1344.Google Scholar
Gubanov, A.P, and Peel, J.S., 2000, Cambrian monoplacophoran molluscs (Class Helcionelloida): American Malacological Bulletin, v. 15(2), p. 139145.Google Scholar
Hamdi, B., Rozanov, A.Y., and Zhuravlev, A.Y., 1995, Latest Middle Cambrian metazoan reef from northern Iran: Geological Magazine, v. 132(4), 3, p. 67373.CrossRefGoogle Scholar
Hartman, W.D., Wendt, J.W., and Wiedenmayer, F., 1980, Living and fossil sponges: notes for a short course, v. 8: Comparative Sedimentology Laboratory, Division of Marine Geology and Geophysics, Rosentiel School of Marine and Atmospheric Science, University of Miami, 274 p.Google Scholar
Hayden, H.H., 1904, The Geology of Spiti with parts of Bashahr and Rupshu: Memoirs of the Geological Survey of India, v. 36, p. 1121.Google Scholar
Hughes, N.C., Peng, S.-C., Bhargava, O.N., Ahulwalia, A.D., Walia, S., Myrow, P.M., and Parcha, S.K., 2005, The Cambrian biostratigraphy of the Tal Group, Lesser Himalaya, India, and early Tsanglangpuan (late early Cambrian) trilobites from the Nigali Dhar syncline: Geological Magazine, v. 142, p. 5780.CrossRefGoogle Scholar
Janussen, D., Steiner, N., and Maoyan, Z., 2002, New well-preserved scleritomes of Chancelloridae from the early Cambrian Yuanshan Formation (Chengjiang, China) and the middle Cambrian Wheeler Shale (Utah, USA) and paleobiological implications: Journal of Paleontology, v. 76, p. 596606.CrossRefGoogle Scholar
Jell, P.A., and Hughes, N.C., 1997, Himalayan Cambrian trilobites: Special Papers in Palaeontology, v. 58, p. 1113.Google Scholar
Jensen, S.R., Palacois, T., and Marti Mus, M., 2010, Revised biochronology of the Lower Cambrian of the Central Iberian zone, southern Iberian Massif, Spain: Geological Magazine, v. 147(5), p. 690703.CrossRefGoogle Scholar
Khomentovsky, V.V., Val’kov, A.K., and Karlova, G.A., 1990, Novye dannye po biostratigrafii perekhodnykh vend-kembrijskikh sloev v bassejne srednego techeniya r. Aldan, Pozdnij dokembrij i rannij paleozoj Sibiri: Voprosy Regional’noj Stratigrafii, p. 357. [in Russian]Google Scholar
Kobayashi, T., 1934, Middle Cambrian fossils from Kashmir: American Journal of Science, v. 27, p. 295302.CrossRefGoogle Scholar
Kouchinsky, A., Bengtson, S., Runnegar, B., Skovsted, C., Steiner, M., and Vendrasco, M., 2012, Chronology of early Cambrian biomineralization: Geological Magazine, v. 149(2), p. 221251.CrossRefGoogle Scholar
Kruse, P.D., 1991, Cambrian fauna of the Top Springs limestone, Georgina basin: The Beagle; Records of the Northern Territory Museum of Arts and Sciences, v. 8, p. 169188.Google Scholar
Kruse, P.D., Laurie, J.R., and Webby, B.D., 2004, Cambrian geology and palaeontology of the Ord Basin: Memoirs of the Association of Australiasian Palaeontologists, v. 30, p. 158.Google Scholar
Kumar, G., Raina, B.K., Bhatt, D.K., and Jangpangi, B.S., 1983, Lower Cambrian body- and trace-fossils from the Tal Formation, Garhwal Synform, Uttar Pradesh, India: Journal of the Palaeontological Society of India, v. 28, p. 106111.Google Scholar
Kumar, G., and Verma, R.N., 1987, Early Cambrian (Tsanglangpuian Stage) trilobites from Vel Nagabal, Anantnag District, Jammu and Kashmir, India: Journal of the Palaeontological Society of India, v. 32, p. 131135.Google Scholar
Kumar, G., Bhatt, D.K., and Raina, B.K., 1987, Skeletal microfauna of Meishucunian and Qiongzhusian (Precambrian-Cambrian boundary) age from the Ganga Valley, Lesser Himalaya, India: Geological Magazine, v. 124, p. 167171.CrossRefGoogle Scholar
Laibl, L., Fatka, O., Cronier, C., and Budil, P., 2014, Early ontogeny of the Cambrian trilobite Sao hirsuta from the Skrye-Tyrovice Basin, Barrandian area, Czech Republic: Bulletin of Geosciences, v. 89, p. 293309.CrossRefGoogle Scholar
Lee, B.S., 2008, Stratigraphic implications of skeletal microfossils from the Cambrian of Korea: a preliminary report: Acta Geologica Sinica, v. 82(6), p. 11521159.CrossRefGoogle Scholar
Lee, D.C., and Chatterton, B.D.E., 2005a, Protaspides of Upper Cambrian Aphelaspis (Ptychopariida, Trilobita) and related species with their taxonomic implications: Palaeontology, v. 48, p. 13511375.CrossRefGoogle Scholar
Lee, D.C., and Chatterton, B.D.E., 2005b, Protaspid ontogeny of Bolaspidella housensis (Order Ptychopariida, Class Trilobita), and other similar Cambrian protaspides: Transactions of the Royal Society of Edinburgh: Earth Sciences, v. 96, p. 2141.CrossRefGoogle Scholar
Lee, H.Y., 1988, Discovery of the early Cambrian small shelly fossils from the Choseon Supergroup at the Kurangni area, Mungyong-Kun, South Korea: Journal of the Paleontological Society of Korea, v. 3, p. 93107.Google Scholar
Luo, H.-L., Jiang, S.-W., Wu, X.-C., Song, X.-L., and Ouyang, L., 1982, [The Sinian-Cambrian boundary in eastern Yunnan, China]: Yunnan, People’s Publishing House, 267 p. [in Chinese with English summary]Google Scholar
Malinky, J.M., and Skovsted, C.B., 2004, Hyoliths and small shelly fossils from the lower Cambrian of north-east Greenland: : Acta Paleontologica Polonica, v. 49, p. 551578.Google Scholar
Maloof, A.C., Porter, S.M., Moore, J.L., Dudás, F.Ö., Bowring, S.A., Higgins, J.A., Fike, D.A., and Eddy, M.P., 2010, The earliest Cambrian record of animals and ocean geochemical change: Geological Society of America Bulletin, v. 122(11–12), p. 17311774.CrossRefGoogle Scholar
Missarzhevsky, V.V., and Mambetov, A.M., 1981, The Stratigraphy and Fauna of the Cambrian-Precambrian Boundary Beds of the Lesser Karatau Range: Trudy Geologii Institut, Akademyia nauk SSSR, v. 326, p. 735738. [in Russian]Google Scholar
Moore, J.L., Li, G., and Porter, S.M., 2014, Chancelloriid sclerites from the Lower Cambrian (Meishucunian) of eastern Yunnan, China, and the early history of the group: Palaeontology, v. 57(4), p. 833878.CrossRefGoogle Scholar
Mostler, H., 1980, Zur Mikrofauna des Unterkambriums in der Haziraformation-Hazara, Pakistan: Naturhistorisches Museum Wien Annalen, v. 83, p. 245257.Google Scholar
Mostler, H., and Mosleh-Yazdi, A., 1976, Neue Poriferen aus oberkambrischen Gesteinen der Milaformation im Elburzgebirge (Iran): Innsbruck, Geologisch-Paläontologische Mitteilungen, v. 5, p. 136.Google Scholar
Myrow, P.M., Snell, K.E., Hughes, N.C., Paulsen, T.S., Heim, N.A., and Parcha, S.K., 2006a, Cambrian depositional history of the Zanskar Valley region of the Indian Himalaya: tectonic implications: Journal of Sedimentary Research, v. 76(2), p. 364381.CrossRefGoogle Scholar
Myrow, P.M., Thompson, K.R., Hughes, N.C., Paulsen, T.S., Sell, B.K., and Parcha, S.K., 2006b, Cambrian stratigraphy and depositional history of the northern Indian Himalaya, Spiti Valley, north-central India: Geological Society of America Bulletin, v. 118(3–4), p. 491510.CrossRefGoogle Scholar
Parkhaev, P.Y., and Demidenko, Y.E., 2010, Zooproblematica and mollusca from the Lower Cambrian Meishucun section (Yunnan, China) and taxonomy and systematics of the Cambrian small shelly fossils of China: Paleontological Journal, v. 44(8), p. 8831161.CrossRefGoogle Scholar
Peel, J.S., 1991a, Functional morphology of the Class Helcionelloida nov., and the early evolution of the Mollusca, in Simonetta, A., and Conway Morris, S., eds., The early evolution of Metazoa and the significance of problematic taxa, Cambridge, Cambridge University Press, p. 157177.Google Scholar
Peel, J.S., 1991b, The classes Tergomya and Helcionelloida, and early molluscan evolution: Bulletin Grønlands Geologiske Undersøgelse, v. 161, p. 1165.Google Scholar
Peng, S.-C., Hughes, N.C., Heim, N.A., Sell, B.K., Zhu, X., Myrow, P.M., and Parcha, S.K., 2009, Cambrian trilobites from the Parahio and Zanskar valleys, Indian Himalaya: Paleontological Society Memoirs (supplement to Journal of Paleontology), v. 71, p. 195.Google Scholar
Peng, S.-C., Uchman, A., Geyer, G., and Hamdi, B., 1999, Trilobites from the Shahmirzad section, Alborz Mountains, Iran: their taxonomy, biostratigraphy and bearing for international correlation: Beringeria, v. 25, p. 366.Google Scholar
Popov, L.E., Holmer, L.E., Hughes, N.C., Ghobadi Pour, M., and Myrow, P.M., 2015, Himalayan Cambrian Brachiopods: Papers in Palaeontology, v. 1(4), p. 345399.CrossRefGoogle Scholar
Porter, S.M., 2004, Closing the phosphatization window: testing for the influence of taphonomic megabias on the pattern of small shelly fossil decline: Palaios, v. 19(2), p. 178183.2.0.CO;2>CrossRefGoogle Scholar
Qian, Y., 1977, Hyolitha and some problematica from the Lower Cambrian Meishucunian Stage in central and southwestern China: Acta Palaeontologica Sinica, v. 16, p. 255275. [in Chinese with English summary]Google Scholar
Qian, Y., and Bengtson, S., 1989, Palaeontology and biostratigraphy of the Early Cambrian Meishucunian Stage in Yunnan Province, South China: Fossils and Strata, v. 24, p. 1156.Google Scholar
Ramakrishnan, M., and Vaidyanadhan, R., 2008, Geology of India, 2 volumes, Bangalore, Geological Society of India, 984 p.Google Scholar
Randell, R.D., Lieberman, B.S., Hasiotis, S.T., and Pope, M.C., 2005, New chancelloriids from the Early Cambrian Sekwi Formation with a comment on chancelloriid affinities: Journal of Paleontology, v. 79, p. 987996.CrossRefGoogle Scholar
Redlich, K.A., 1899, The Cambrian fauna of the eastern Salt-Range: Palaeontologia Indica New Series, v. 1(1), p. 113.Google Scholar
Reed, F.R.C., 1910, The Cambrian fossils of Spiti: Palaeontologica Indica Series 1, v. 57, p. 170.Google Scholar
Reed, F.R.C., 1934, Cambrian and Ordovician fossils from Kashmir: Palaeontologica Indica, v. 21(2), p. 138.Google Scholar
Rozanov, A.Yu., Missarzhevsky, V.V., Volkova, N.A., Voronova, L.G., Krylov, I.N., Keller, B.M., Korolyuk, I.K., Lendzion, K., Mikhnyar, R., Pykhova, N.G., and Sidorov, A.D., 1969, Tommotskiu jarus i problema nizhney granisty kembriya: Trudy Geoligske Institut Leningrad, v. 206, p. 1379. [in Russian]Google Scholar
Schindewolf, O.H., 1955, Die Körperfossilien des Unterkambriums: Akademie der Wissenschaften und der Literatur, Mainz Mathematisch-naturwissenschaflichen Klasse Abhandlungen, v. 10, p. 281341.Google Scholar
Sdzuy, K., 1969, Unter- und mittelkambrische Porifera (Chancelloriida und Hexactinellida): Paläontologische Zeitschrift, v. 43, p. 115147.CrossRefGoogle Scholar
Shah, S.K., and Raina, A.K., 1990, Middle-Late Cambrian transition in Kashmir and Spiti Himalaya: Memoirs of the Geological Society of India, v. 16, p. 4150.Google Scholar
Shah, S.K., Parcha, S.K., and Raina, A.K., 1991, Late Cambrian trilobites from Himalaya: Journal of the Palaeontological Society of India, v. 36, p. 89107.Google Scholar
Singh, B.P., Bhargava, O.N., Juyal, K.P., Negi, R.S., Virmani, N., Sharma, C.A., and Gill, A., 2015, Skeletal microfauna from the Cambrian Series 2 (Stage 4) Kunzum La Formation, Parahio valley, Spiti region (Tethyan Himalaya), India: Current Science, v. 109(12), p. 21912195.Google Scholar
Skovsted, C.B., and Peel, J.S., 2007, Small shelly fossils from the argillaceous facies of the Lower Cambrian Forteau Formation of western Newfoundland: Acta Palaeontologica Polonica, v. 52(4), p. 729748.Google Scholar
Sollas, W.J., 1888, Report on the Tetractinellida collected by H.M.S. Challenger during the years 1873–1876: Report on Scientific Researches of H.M.S. Challenger, v. 25, 458+166 p.Google Scholar
Swinnerton, H.H., 1915, Suggestions for a revised classification of trilobites: Geological Magazine (New Series), v. 6(2), p. 487496, 538–545.CrossRefGoogle Scholar
Tiwari, M., 1989, Discovery of pre-trilobite small shelly fossils and position of Precambrian-Cambrian boundary in Tethyan sequence or northwestern Kashmir: Current Science, v. 58, p. 839843.Google Scholar
Vasil’eva, N.I., 1985, K sistematike otryada Chancelloriida Walcott, 1920 (incertae sedis) iz nizhnekembriyskikh otlozheniy vostoka Sibirskoy platform, in Sokolov, B.S., and Zhuravleva, I.T., eds., Problematiki pozdnego dokembriya i paleozoya: Akademiya Nauk SSSR, Sibirskoe Otdelenie, Trudy Instituta Geologii i Geofiziki, 632, p. 115–126. [in Russian]Google Scholar
Vasil’eva, N.I., and Sayutina, T.A., 1988, The morphological diversity of chancelloriid scleritis: Trudy Instituta geologii i geofiziki (Novobisibirsk), v. 720, p. 190198.Google Scholar
Vasil’eva, N. I., and Sayutina, T.A., 1993, Novye rodovye i vidovoe nazvanie rannekembriyskikh skleritov khantselloriid: Paleontological Zhornal, v. 1993(1), p. 113114. [in Russian]Google Scholar
Von Siebold, C.T., 1848, Lehrbuch der vergleichenden Anatomie der Wirbellosen Thiere. Erster Theil, in Von Siebold, C.T., and Stannius, H., eds., Lehrbuch der vergleichenden Anatomie, Berlin, Verlag von Veit and Comp, 679 p.Google Scholar
Vostokova, V.A., 1962, Kembrijskie gastropody Sibirskoj platformy i Tajmyra: Trudy Nauchno Issledovatel’skogo Instituta Geologii Arktiki, v. 28, p. 5174.Google Scholar
Waagen, W., 1882–1885, Salt range fossils. I. Productus limestone fossils: Palaeontologica Indica Series 13, v. 1(4), p. 329770.Google Scholar
Waagen, W., 1891, Salt range fossils: Palaeontologia Indica Series 13, v. 4, p. 89242.Google Scholar
Walch, J.E.I., 1771, Die Naturgeschichte der Versteinerungen Dritter Theil, Nuremberg, Paul Jonathan Felstecher, 235 p.Google Scholar
Walcott, C.D., 1920, Middle Cambrian Spongiae: Smithsonian Miscellaneous Collections, v. 67, p. 261364.Google Scholar
Wenz, W., 1938–1943, Gastropoda. Allgemeiner Teil und Prosobranchia. Handbuch der Palaozoologie, Berlin, Gebruder Bornträger, 1506 p.Google Scholar
Wrona, R., 2003, Early Cambrian molluscs from glacial erratics of King George Island, west Antarctica: Polish Polar Research, v. 24, p. 181216.Google Scholar
Xing, Y.-S., Ding, Q.-X., Luo, H.-L., He, T.-G., and Wang, Y.-G., 1984, The Sinian-Cambrian boundary of China and its related problems: Geological Magazine, v. 121(3), p. 155170.Google Scholar
Yu, W., 1974, Cambrian Gastropoda, in Handbook of the Stratigraphy and Paleontology of Southwest China, Beijing, Science Press, p. 111113. [in Chinese]Google Scholar
Yu, W., 1979, Earliest Cambrian Monoplacophorans and Gastropods from Western Hubei with their biostratigraphical significance: Acta Palaeontologica Sinica, v. 18(3), p. 233270. [in Chinese]Google Scholar
Yu, W., 1987, Yangtze micromolluscan fauna in Yangtze region of China with notes on Precambrian-Cambrian boundary, in Stratigraphy and Palaeontology of Systemic Boundaries in China–Precambrian–Cambrian Boundary 1, Nanjing, Nanjing University Publishing House, p. 19344.Google Scholar
Zhang, X.-G., and Pratt, B.R., 1999, Early Cambrian trilobite larvae and ontogeny of Ichangia ichangensis Chang 1957 (Protolenidae) from Henan, China: Journal of Paleontology, v. 73, p. 117128.CrossRefGoogle Scholar
Zhao, Y.-L., Zhu, M.-Y., Babcock, L.E., and Peng, J., 2011, The Kaili Biota: Marine Organisms from 508 Million Years Ago, Guiyang, Guizhou Science and Technology Press, 251 p.Google Scholar
Zhou, B., and Xiao, L., 1984, (Early Cambrian monoplacophorans and gastropods from Huainan and Huoqui counties, Anhui Province): Professional Papers of Stratigraphy and Palaeontology, Chinese Academy of Geological Sciences, v. 13, p. 125140. [in Chinese]Google Scholar
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