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CATALOGUE OF THE GEADEPHAGA (COLEOPTERA: TRACHYPACHIDAE, RHYSODIDAE, CARABIDAE INCLUDING CICINDELINI) OF AMERICA NORTH OF MEXICO

Published online by Cambridge University Press:  31 May 2012

Y. Bousquet  and
A. Larochelle
Y. Bousquet
Affiliation:
Centre for Land and Biological Resources Research, Biological Resources Division, Research Branch, Agriculture Canada, Central Experimental Farm, Ottawa, Ontario, Canada KIA 0C6.
A. Larochelle
Affiliation:
Native Plants and Animals Division, Landcare Research New Zealand Ltd.. Private Bag 92170, Auckland, New Zealand
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Abstract

All species-group names of Trachypachidae, Rhysodidae, and Carabidae (including cicindelincs) correctly recorded from America north of Mexico are catalogued with state and province records. Valid names are listed with the author(s), date of publication, and page citation in their current and original combinations while all synonyms are provided in their original combinations. Genus-group names are recorded with the author(s), date of publication, page citation, type species, and kind of type species fixation. Species groups were preferred to subgenera but subscneric names are also listed.The following nomenclatural changes are proposed and discussed: Bembidion neocoerulescens Bousquet, new replacement name for B. coerulescens Van Dyke, 1925; Chlaenius circumcinctus Say, 1830 for C. perplexus Dejean, 1831; Cyclotrachelus dejeanellus (Csiki, 1930) for C. morio (Dejean, 1828); Cyclotrachelus freitagi Bousquet, new replacement name for C. obsoletus (Say, 1830); Dyschirius aeneolus LeConte, 1850 for D. frigidus Mannerheim, 1853; Harpalus laevipes Zetterstedt, 1828 for H. quadripunctatus Dejean, 1829; Harpalus providens Casey, 1914 for H. viduus LeConte, 1865; Harpalus reversus Casey, 1924 for H. funerarius Csiki, 1932; Notiophilus sierranus Casey, 1920 for N. obscurus Fall, 1901; Pseudamara Lindroth, 1968 for Disamara Lindroth, 1976; Pterostichus trinarius (Casey, 1918) for P. ohionis Csiki, 1930; Stenolophus carbo Bousquet, new replacement name for S. carbonarius (Dejean, 1829).Thirty-six new synonyms are established and seven, considered as questionable, are confirmed. They are (with the valid names in parentheses): Agonothorax planipennis Motschulsky, 1850 (= ? Agonum affine Kirby, 1837); Platynus variolatus LeConte, 1851 (= Agonum limbatum Motschulsky, 1845); Agonum nitidum Harris, 1869 (= ? Agonum melanarium Dejean, 1828); Amerinus fuscicornis Casey, 1914 and A. longipennis Casey, 1914 (= Amerinus linearis (LeConte, 1863)); Apristus fuscipennis Motschulsky, 1864 (= Apristus latens LeConte, 1848); Batenus aeneolus Motschulsky, 1865 (= Agonum exaratum (Mannerheim, 1853)); Brachystylus curtipennis Motschulsky, 1859 (= Pterostichus congestus (Ménétriés, 1843)); Brachystylus parallelus Motschulsky, 1859 (= ? Pterostichus californicus (Dejean, 1828)); Cratacanthus cephalotes Casey, 1914, C. subovalis Casey, 1914, and C. texanus Casey, 1884 (= Cratacanthus dubius (Palisot de Beauvois, 1811)); Cymindis comma T.W. Harris, 1869 (= ? Cymindis limbatus Dejean, 1831); Feronia praetermissa Chaudoir, 1868 (= Pterostichus commutabilis (Motschulsky, 1866)); Galerita angusticeps Casey, 1920 (= Galerita janus (Fabricius, 1792)); Gonoderus cordicollis Motschulsky 1859 (= Pterostichus tristis (Dejean, 1828)); Anisodactylus alternans LeConte, 1851 (= Anisodactylus alternans (Motschulsky, 1845)); Hypherpes spissitarsis Casey, 1918 (= Pterostichus tarsalis LeConte, 1873); Lebia brunnicollis Motschulsky, 1864 (= Lebia lobulata LeConte, 1863); Lebia subfigurata Motschulsky, 1864 and L. sublimbata Motschulsky, 1864 (= Lebia analis Dejean, 1825); Lophoglossus bispiculatus Casey, 1913 and L. illini Casey, 1913 (= Lophoglossus scrutator (LeConte, 1848)); Platysma leconteianum Lutshnik, 1922 (= Pterostichus commutabilis (Motschulsky, 1866)); Loxandrus iris Motschulsky, 1866(= Loxandrus rectus (Say, 1823)); Masoreus americanus Motschulsky, 1864 (= Stenolophus rotundicollis (Haldeman, 1843)); Notaphus laterimaculatus Motschulsky, 1859 (= Bembidion approximatum (LeConte, 1852)); Notiophilus cribrilaterus Motschulsky, 1864 (= Notiophilus novemstriatus LeConte, 1848); Omaseus brevibasis Casey, 1924 (= Pterostichus luctuosus (Dejean, 1828)); Notaphus incertus Motschulsky, 1845 (= Bembidion breve (Motschulsky, 1845)); Peryphus concolor Motschulsky, 1850 (= Bembidion platynoides Hayward, 1897); Peryphus erosus Motschulsky, 1850 (= Bembidion transversale Dejean, 1831); Peryphus subinflatus Motschulsky, 1859 (= Bembidion petrosum petrosum Gebler, 1833); Planesus fuscicollis Motschulsky, 1865 and P. laevigatas Motschulsky, 1865 (= Cymindis platicollis (Say, 1823)); Poecilus pimalis Casey, 1913 (= Poecilus diplophryus Chaudoir, 1876); Pterostichus arizonicus Schaeffer, 1910 (= Ophryogaster flohri Bates, 1882); Pterostichus sequoiarum Casey, 1913 (= Pterostichus tarsalis LeConte, 1873); Scaphinotus grandis Gistel, 1857 (= ? Scaphinotus unicolor unicolor (Fabricius, 1787)); Stenocrepis chalcas Bates, 1882 and S. chalcochrous Chaudoir, 1883 (= Stenocrepis texana (LeConte, 1863)); Stenolophus humeralis Motschulsky, 1864 (= Stenolophus plebejus Dejean, 1829); and Stenolophus laticollis Motschulsky, 1864 (= Stenolophus ochropezus (Say, 1823)).Olisthopus iterans Casey, 1913 and Pterostichus illustris LeConte, 1851, listed as junior synonyms of O. parmatus (Say, 1823) and P. congestus (Ménétriés, 1843), respectively, are considered in the present work as valid species.The type species (listed in parentheses) of the following 14 genus-group taxa are designated for the first time: Circinalidia Casey, 1920 (Agonum aeruginosum Dejean, 1828); Evolenes LeConte, 1853 (Oodes exaratus Dejean, 1831); Leucagonum Casey, 1920 (Agonum maculicolle Dejean, 1828); Megaliridia Casey, 1920 (Cychrus viduus Dejean, 1826); Megalostylus Chaudoir, 1843 (Feronia lucidula Dejean, 1828 = Feronia recta Say, 1823); Micragra Chaudoir, 1872 (Micragra lissonota Chaudoir, 1872); Onota Chaudoir, 1872 (Onota bicolor Chaudoir, 1872); Oodiellus Chaudoir, 1882 (Oodiellus mexicanus Chaudoir, 1882 = Anatrichis alutacea Bates, 1882); Oxydrepanus Putzeys, 1866 (Dyschirius rufus Putzeys, 1846); Paranchomenus Casey, 1920 (Platynus stygicus LeConte, 1854 = Anchomenus mannerheimii Dejean, 1828); Pemphus Motschulsky, 1866 (Cychrus velutinus Ménétriés, 1843); Peronoscelis Chaudoir, 1872 (Tetragonoderus figuratus Dejean, 1831); Rhombodera Reiche, 1842 (Rhombodera virgata Reiche, 1842 = Lebia trivittata Dejean, 1831); and Stenous Chaudoir, 1857 (Oodes cupreus Chaudoir, 1843).Two new family-group names are proposed, Cnemalobini (= Cnemacanthini of authors) based on Cnemalobus Guérin-Méneville, 1839 and Loxandrini based on Loxandrus LeConte, 1852.The work also includes a synopsis of all extant world carabid tribes, a bibliography of all original descriptions, a full taxonomic index, and, as appendices, lists of nomina nuda and unjustified emendations, and annotated lists of species incorrectly or doubtfully recorded from America north of Mexico and of new North American records.

Résumé

Ce catalogue comprend tous les noms de rang groupe-espèce des familles Trachypachidae, Rhysodidae, et Carabidae (les cicindelines inclus) correctement recensés de l'Amérique, au nord du Mexique ainsi que leurs mentions d'états et de provinces. On cite les noms valides dans leurs combinaisons actuelle et originale avec l'auteur(s), la date et la page de publication; on mentionne les synonymes dans leur combinaison originale seulement. Pour les noms de rang groupe-genre on indique l'auteur(s), la date et la page de publication, l'espèce-type, et le type de fixation de l'espèce-type. On a préféré le concept de groupe d'espèces à celui de sous-genre mais les noms sous-génériques font également l'objet de mention.

On propose et discute les changements de nomenclature suivants: Bembidion neocoerulescens Bousquet, nouveau nom de remplacement pour B. coerulescens Van Dyke, 1925; Chlaenius circumcinctus Say, 1830 pour C. perplexus Dejean, 1831; Cyclotrachelus dejeanellus (Csiki, 1930) pour C. morio (Dejean, 1828); Cyclotrachelus freitagi Bousquet, nouveau nom de remplacement pour C. obsoletus (Say, 1830); Dyschirius aeneolus LeConte, 1850 pour D. frigidus Mannerheim, 1853; Harpalus laevipes Zetterstedt, 1828 pour H. quadripunctatus Dejean, 1829; Harpalus providens Casey, 1914 pour H. viduus LeConte, 1865; Harpalus reversus Casey, 1924 pour H. funerarius Csiki, 1932; Notiophilus sierranus Casey, 1920 pour N. obscurus Fall, 1901; Pseudamara Lindroth, 1968 pour Disamara Lindroth, 1976; Pterostichus trinarius (Casey, 1918) pour P. ohionis Csiki, 1930; Stenolophus carbo Bousquet, nouveau nom de remplacement pour S. carbonarius (Dejean, 1829).

On établit 36 nouveaux synonymes et sept autres, considérés comme douteux, sont confirmés. Ces synonymes sont (avec le nom valide indiqué entre parenthèses): Agonothorax planipennis Motschulsky, 1850(= ? Agonum affine Kirby, 1837); Platynus variolatus LeConte, 1851 (= Agonum limbatum Motschulsky, 1845); Agonum nitidum Harris, 1869 (= ? Agonum melanarium Dejean, 1828); Amerinus fuscicornis Casey, 1914 et A. longipennis Casey, 1914 (= Amerinus linearis (LeConte, 1863)); Apristus fuscipennis Motschulsky, 1864 (= Apristus latens LeConte, 1848); Batenus aeneolus Motschulsky, 1865 (= Agonum exaratum (Mannerheim, 1853)); Brachystylus curtipennis Motschulsky, 1859 (= Pterostichus congestus (Ménétriés, 1843)); Brachystylus parallelus Motschulsky, 1859 (= ? Pterostichus californicus (Dejean, 1828)); Cratacanthus cephalotes Casey, 1914, C. subovalis Casey, 1914 et C. texanus Casey, 1884 (= Cratacanthus dubius (Palisot de Beauvois, 1811)); Cymindis comma T.W. Harris, 1869 (= Cymindis limbatus Dejean, 1831); Feronia praetermissa Chaudoir, 1868 (= Pterostichus commutabilis (Motschulsky, 1866)); Galerita angusticeps Casey, 1920 (= Galerita janus (Fabricius, 1792)); Gonoderus cordicollis Motschulsky, 1859 (= Pterostichus tristis (Dejean, 1828)); Anisodactylus alternans LeConte, 1851 (= Anisodactylus alternans (Motschulsky, 1845)); Hypherpes spissitarsis Casey, 1918 (= Pterostichus tarsalis LeConte, 1873); Lebia brunnicollis Motschulsky, 1864 (= Lebia lobulata LeConte, 1863); Lebia subfigurata Motschulsky, 1864 et L. sublimbata Motschulsky, 1864 (= Lebia analis Dejean, 1825); Lophoglossus bispiculatus Casey, 1913 et L. illini Casey, 1913 (= Lophoglossus scrutator (LeConte, 1848)); Platysma leconteianum Lutshnik, 1922 (= Pterostichus commutabilis (Motschulsky, 1866)); Loxandrus iris Motschulsky, 1866 (= Loxandrus rectus (Say, 1823)); Masoreus americanus Motschulsky, 1864 (= Stenolophus rotundicollis (Haldeman, 1843)); Notaphus laterimaculatus Motschulsky, 1859 (= Bembidion approximatum (LeConte, 1852)); Notiophilus cribrilaterus Motschulsky, 1864 (= Notiophilus novemstriatus LeConte, 1848); Omaseus brevibasis Casey, 1924 (= Pterostichus luctuosus (Dejean, 1828)); Notaphus incertus Motschulsky, 1845 (= Bembidion breve (Motschulsky, 1845)); Peryphus concolor Motschulsky, 1850 (= Bembidion platynoides Hayward, 1897); Peryphus erosus Motschulsky, 1850 (= Bembidion transversale Dejean, 1831); Peryphus subinflatus Motschulsky, 1859 (= Bembidion petrosum petrosum Gebler, 1833); Planesus fuscicollis Motschulsky, 1865 et P. laevigatus Motschulsky, 1865 (= Cymindis platicollis (Say, 1823)); Poecilus pimalis Casey, 1913 (= Poecilus diplophryus Chaudoir, 1876); Pterostichus arizonicus Schaeffer, 1910(= Ophryogaster flohri Bates, 1882); Pterostichus sequoiarum Casey, 1913 (= Pterostichus tarsalis LeConte, 1873); Scaphinotus grandis Gistel, 1857 (= ? Scaphinotus unicolor unicolor (Fabricius, 1787)); Stenocrepis chalcas Bates, 1882 et S. chalcochrous Chaudoir, 1883 (= Stenocrepis texana (LeConte, 1863)); Stenolophus humeralis Motschulsky, 1864 (= Stenolophus plebejus Dejean, 1829); et Stenolophus laticollis Motschulsky, 1864 (= Stenolophus ochropezus (Say, 1823)).

Olisthopus iterans Casey, 1913 et Pterostichus illustris LeConte, 1851, listés comme synonymes récents de O. parmatus (Say, 1823) et P. congestus (Ménétriés, 1843) respectivement, sont considérés dans ce catalogue comme espèces valides.

On désigne pour la première fois l'espèce-type (citée entre parenthèses) des 14 taxa suivants : Circinalidia Casey, 1920 (Agonum aeruginosum Dejean, 1828); Evolenes LeConte, 1853 (Oodes exaratus Dejean, 1831); Leucagonum Casey, 1920 (Agonum maculicolle Dejean, 1828); Megaliridia Casey, 1920 (Cychrus viduus Dejean, 1826); Megalostylus Chaudoir, 1843 (Feronia lucidula Dejean, 1828 = Feronia recta Say, 1823); Micragra Chaudoir, 1872 (Micragra lissonota Chaudoir, 1872); Onota Chaudoir, 1872 (Onota bicolor Chaudoir, 1872); Oodiellus Chaudoir, 1882 (Oodiellus mexicanus Chaudoir, 1882 = Anatrichis alutacea Bates, 1882); Oxydrepanus Putzeys, 1866 (Dyschirius rufus Putzeys, 1846); Paranchomenus Casey, 1920 (Platynus stygicus LeConte, 1854 = Anchomenus mannerheimii Dejean, 1828); Pemphus Motschulsky, 1866 (Cychrus velutinus Ménétriés, 1843); Peronoscelis Chaudoir, 1872 (Tetragonoderus figuratus Dejean, 1831); Rhombodera Reiche, 1842 (Rhombodera virgata Reiche, 1842 = Lebia trivittata Dejean, 1831); et Stenous Chaudoir, 1857 (Oodes cupreus Chaudoir, 1843).

On propose deux nouveaux noms du rang de groupe-famille, Cnemalobini (= Cnemacanthini des auteurs) fondé sur Cnemalobus Guérin-Méneville, 1839 et Loxandrini fondé sur Loxandrus LeConte, 1852.

L'ouvrage comprend aussi un synopsis de toutes les tribus de Carabidae existantes, une bibliographie de toutes les descriptions originales, un index taxonom

Type
Research Article
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The Memoirs of the Entomological Society of Canada , Volume 125 , Supplement S167 , 1993 , pp. 3 - 397
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Copyright © Entomological Society of Canada 1993

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