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  • R. R. Mill (a1)


The species of Podocarpus L’Hér. ex Pers. (Podocarpaceae) occurring in the Central America and Northern Mexico Bioregions are revised. Four species (Podocarpus costaricensis de Laub., P. guatemalensis Standl., P. matudae Lundell, P. oleifolius D.Don) occur in these bioregions as well as three infraspecific taxa, that are here all treated as subspecies [Podocarpus matudae subsp. matudae, P. matudae subsp. jaliscanus (de Laub. & Silba) Silba, P. oleifolius subsp. costaricensis (J.Buchholz & N.E.Gray) Silba]. A fifth species, Podocarpus magnifolius J.Buchholz & N.E.Gray, may also be present in Panama but this requires verification; a brief account is provided. Podocarpus monteverdeensis de Laub. is considered a synonym of P. oleifolius subsp. costaricensis, the concept of which is amplified to include all Central American material of P. oleifolius. Several previously recognised infraspecific taxa within both Podocarpus guatemalensis and P. matudae are reduced to synonymy. Within Podocarpus matudae, P. matudae subsp. matudae is regarded as including subsp. macrocarpus and subsp. reichei but P. matudae subsp. jaliscanus is regarded as a distinct, second subspecies disjunct in westernmost Mexico. Podocarpus costaricensis, P. matudae (both subspecies) and P. oleifolius subsp. costaricensis are endemic to these bioregions. A key is provided, all definitely recorded species are illustrated and the distributions of all definitely recorded taxa are mapped. The distributions are discussed in relation to the geology and geological history of the region as well as altitude and climate. New IUCN conservation assessments are proposed for Podocarpus matudae subsp. jaliscanus, P. matudae subsp. matudae and P. oleifolius subsp. costaricensis while details of the current assessments for the remaining taxa (including Podocarpus matudae as a whole) are given. Two appendices list all accepted names and synonyms, and give a list of exsiccatae.



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This and the previous paper (Mill, 2015) are dedicated to the late Darian Stark Schilling (1980–2011).



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Alcántara Ayala, O. & Luna Vega, I. (2001). Análisis florístico de dos áreas con bosque mesófilo de montaña en el Estado de Hidalgo, México: Eloxochitlán y Tlahuelompa. Acta Bot. Mex. 54: 5187.
Alvarado, G. E. (1985). Geomorfología y petrografía de algunas provincias volcánicas de Costa Rica. Bol. Vulcanol. (Heredia) 14: 2428.
Arguedas, M. (2007). Plagas y enfermedades forestales en Costa Rica. Kurú (Rev. Forest.) 4 (11 & 12 especial): i–vi, 169.
Arriaga, L. (1988). Gap dynamics of a tropical cloud forest in Northeastern Mexico. Biotropica 20: 178184.
Arriaga, L. (2000a). Gap-building-phase regeneration in a tropical montane cloud forest of north-eastern Mexico. J. Trop. Ecol. 16: 535562.
Arriaga, L. (2000b). Types and causes of tree mortality in a tropical montane cloud forest of Tamaulipas, Mexico. J. Trop. Ecol. 16: 623636.
Azéma, J. & Tournon, J. (1982). The Guatemalan margin, the Nicoya Complex, and the origin of the Caribbean Plate. In: Audouin, J., von Huene, R., Azema, J., Coulbourn, W. T., Cowan, S. S., Curiale, J. A., Dengo, C. A., Faas, R. W., Harrison, W. E., Hesse, R., Ladd, J. W., Muzylov, N., Shiki, T., Thomson, P. R., Westberg, M. J. & Orlofsky, S. (eds) Init. Rep. Deep Sea Drill. Proj. 67: 739745. Washington: US Government Printing Office.
Bachman, S., Farjon, A., Gardner, M., Thomas, P., Luscombe, D. & Reynolds, C. (2007a). Podocarpus costaricensis. In: IUCN (2010). IUCN Red List of Threatened Species, Version 2011.2., downloaded 10 February 2012.
Bachman, S., Farjon, A., Gardner, M., Thomas, P., Luscombe, D. & Reynolds, C. (2007b). Podocarpus guatemalensis. In: IUCN (2010). IUCN Red List of Threatened Species, Version 2011.2., downloaded 10 February 2012.
Bachman, S., Farjon, A., Gardner, M., Thomas, P., Luscombe, D. & Reynolds, C. (2007c). Podocarpus matudae. In: IUCN (2010). IUCN Red List of Threatened Species, Version 2010.3., downloaded 30 September 2010.
Bacon, C. D., Mora, A., Wagner, W. L. & Jaramillo, C. A. (2013). Testing geological models of evolution of the Isthmus of Panama in a phylogenetic framework. Bot. J. Linn. Soc. 171: 287300.
Balick, M. J., Nee, M. H. & Atha, D. E. (2000). Checklist of the Vascular Plants of Belize with Common Names and Uses. Bronx, NY: The New York Botanical Garden Press. (Being Mem. New York Bot. Gard. 85).
Bannister, P. & Neuner, G. (2001). Frost resistance and the distribution of conifers. In: Bigras, F. J. & Colombo, S. J. (eds) Conifer Cold Hardiness, pp. 322. Dordrecht: Kluwer Academic Publishers.
Barreda de Hurtado, L., Cabrera, R. O., Galtán, J. M. & González, J. (1999). Plan de Desarrollo sostenible de la Zona Fronteriza Guatemala-Honduras en Torno al Golfo de Honduras. Factibilidad. Proyecto: Manejo ecoturistico en areas protegidas. Area Izabal, Guatemala. Informe de Consultoría. Coronado: IICA (Instituto Interamericano de Cooperación para la Agricultura).
Barrier, E., Velasquillo, L., Chavez, M. & Gaulon, R. (1998). Neotectonic evolution of the Isthmus of Tehuantepec (southeastern Mexico). Tectonophysics 287: 7796.
Bartlett, H. H. (1935). A method of procedure for field work in tropical American phytogeography based upon a botanical reconnaissance in parts of British Honduras and the Peten forest of Guatemala. Publ. Carnegie Inst. Washington 461: 125.
Bauch, J., Koch, G., Puls, J., Schwarz, T. & Voiß, S. (2006a). Wood characteristics of Podocarpus oleifolius var. macrostachyus (Parl.) Buchholz and Gray native to Costa Rica: their significance for wood utilization. Wood Sci. Technol. 40: 2638.
Bauch, J., Quiros, L., Noldt, G. & Schmidt, P. (2006b). Study on the wood anatomy, annual wood increment and intra-annual growth dynamics of Podocarpus oleifolius var. macrostachyus from Costa Rica. J. Appl. Bot. Food Qual. 80: 1924.
Baumgartner, P. O., Flores, K., Bandini, A. N., Girault, F. & Cruz, D. (2008). Upper Triassic to Cretaceous Radiolaria from Nicaragua and northern Costa Rica: the Mesquito Composite Oceanic Terrane. Ofioliti 33: 119.
Benny, G. L., Samuelson, D. A. & Kimbrough, J. W. (1985a). Studies on the Coryneliales. II. Taxa parasitic on Podocarpaceae: Corynelia. Bot. Gaz. 146: 238251.
Benny, G. L., Samuelson, D. A. & Kimbrough, J. W. (1985b). Studies on the Coryneliales. III. Taxa parasitic on Podocarpaceae: Lagenulopsis and Tripospora. Bot. Gaz. 146: 431436.
Biffin, E., Conran, J. G. & Lowe, A. J. (2011). Podocarp evolution: A molecular phylogenetic perspective. In: Turner, B. L. & Cernusak, L. A. (eds) Ecology of the Podocarpaceae in Tropical Forests, pp. 120. Smithsonian Contributions to Botany, No. 95. Washington, DC: Smithsonian Institution Scholarly Press.
Biffin, E., Brodribb, T. J., Hill, R. S., Thomas, P. & Lowe, A. J. (2012). Leaf evolution in Southern Hemisphere conifers tracks the angiosperm ecological radiation. Proc. Roy. Soc. B 279: 341348.
Bird, P. (2003). An updated digital model of plate boundaries. Geochem., Geophys., Geosyst. 4(3): article 1027. doi:10.1029/2001GC000252, downloaded 9 October 2013.
Blount, D. N. & Moore, C. H. Jr. (1969). Depositional and non-depositional carbonate breccias, Chiantla Quadrangle, Guatemala. Geol. Soc. Amer. Bull. 80: 429442.
Brako, L. & Zarucchi, J. L. (1993). Catalogue of the flowering plants and gymnosperms of Peru. Monogr. Syst. Bot. Missouri Bot. Gard. 45. St. Louis, MO: Missouri Botanical Garden.
Bridges, E. M. (1990). World Geomorphology. Cambridge, UK: Cambridge University Press.
Bridgewater, S. G. M., Harris, D. J., Whitefoord, C., Monro, A. K., Penn, M. G., Sutton, D. A., Sayer, B., Adams, B., Balick, M. J., Atha, D. H., Solomon, J. & Holst, B. K. (2006). A preliminary checklist of the vascular plants of the Chiquibul Forest, Belize. Edinburgh J. Bot. 63: 269321.
Brodribb, T. (2011). A functional analysis of podocarp ecology. Smithsonian Contrib. Bot. 95: 165173.
Brummitt, R. K. (2001). World Geographical Scheme for Recording Plant Distributions Edition 2. Plant Taxonomic Database Standards No. 2. Pittsburgh: Hunt Institute for Botanical Documentation, Carnegie Mellon University, for the International Working Group on Taxonomic Databases for Plant Sciences (TDWG).
Buchholz, J. T. & Gray, N. E. (1948a). A taxonomic revision of Podocarpus. I. The sections of the genus and their subdivisions with special reference to leaf anatomy. J. Arnold Arbor. 29: 4963.
Buchholz, J. T. & Gray, N. E. (1948b). A taxonomic revision of Podocarpus IV. The American species of section Eupodocarpus, Subsections C and D. J. Arnold Arbor. 29: 123151.
Buchs, D. M., Arculus, R. J., Baumgartner, P. O., Baumgartner-Mora, C. & Ulianov, A. (2010). Late Cretaceous arc development on the SW margin of the Caribbean Plate: Insights from the Golfito, Costa Rica, and Azuero, Panama, complexes. Geochem., Geophys., Geosyst. 11(7): Q07S24. 35 pp. doi:10.1029/2009GC002901, downloaded 9 October 2013.
Buchs, D. M., Arculus, R. J., Baumgartner, P. O. & Ulianov, A. (2011). Oceanic intraplate volcanoes exposed: Example from seamounts accreted in Panama. Geology 39: 335338.
Bueno, J., Álvarez, F. & Santiago, S. (eds) (2005). Biodiversidad del Estado de Tabasco. Ciudad Universitaria, México City: CONABIO/Instituto de Biología, UNAM.
Carlson, M. C. (1954). Floral elements of the Pine-Oak-Liquidambar forest of Montebello, Chiapas, Mexico. Bull. Torrey Bot. Club 81: 387399.
Carrasquilla, L. G. (1997). CPD Site MA19, Cerro Azul-Cerro Jefe Region, Panama. In: Davis, S. D., Heywood, V. H., Herrera-MacBryde, O., Villa-Lobos, J. & Hamilton, A. (eds) Centres of Plant Diversity: A Guide and Strategy for their Conservation, Volume 3: The Americas, pp. 221225. Cambridge, UK: IUCN Publications Unit. Also available at, accessed 27 September, 31 October 2013.
Castillo, M. L. & Reyes-Castillo, P. (2003). Los Passalidae: coléopteros tropicales degradadores de troncos de árboles muertos. In: Álvarez-Sánchez, J. & Naranjo-García, E. (eds) Ecología del suelo en la selva tropical húmido de México, pp. 237260. Xalapa: Instituto de Ecologia, A.C., Instituto de Biología y Facultad de Ciencias, UNAM.
Castillo Lemus, N. A. (2005). Estudio de identificación y caracterización preliminar de corredores biológicos con conectividad al biotopo Chocón Machacas. Guatemala City: CECON, Universidad de San Carlos., downloaded 4 November 2013.
Cavelier, J., Solis, D. & Jaramillo, M. A. (1996). Fog interception in montane forests across the Central Cordillera of Panama. J. Trop. Ecol. 12: 357369.
Cayuela, L., Golicher, D. J. & Rey-Benayas, J. M. (2006a). The extent, distribution, and fragmentation of vanishing montane cloud forest in the highlands of Chiapas, Mexico. Biotropica 38: 544554.
Cayuela, L., Rey Benayas, J. M. & Echevarría, C. (2006b). Clearance and fragmentation of tropical montane forests in the Highlands of Chiapas, Mexico (1975–2000). Forest Ecol. Managem. 226: 208218.
Clemons, R. E. & Burkart, B. (1971 [‘1969’]). Stratigraphy of northwestern Guatemala. Bol. Soc. Geol. Mex. 32: 143158.
Coates, A. G. & Obando, J. A. (1996). The geological evolution of the Central American Isthmus. In: Jackson, J. B. C., Budd, A. F. & Coates, A. G. (eds) Evolution and Environment in Tropical America, pp. 2156. Chicago: University of Chicago Press.
Coates, A. G., Collins, L. S., Aubry, M.-P. & Berggren, W. A. (2004). The geology of the Darien, Panama, and the late Miocene-Pliocene collision of the Panama arc with northwestern South America. Geol. Soc. Amer. Bull. 116: 13271344.
Coates, A. G., McNeill, D. F., Aubry, M. P., Berggren, W. A. & Collins, L. S. (2005). An introduction to the geology of the Bocas del Toro Archipelago, Panama. Caribbean J. Sci. 41: 374391.
Colinvaux, P. A., De Oliveira, P. E. & Bush, M. B. (2000). Amazonian and neotropical plant communities on glacial time-scales: The failure of the aridity and refuge hypotheses. Quatern. Sci. Rev. 19: 141169.
Condit, R., Pérez, R. & Daguerre, N. (2011). Trees of Panama and Costa Rica. Princeton, NJ: Princeton University Press.
Contreras-Medina, R., Luna-Vega, I. & Alcántara-Ayala, O. (2006). La familia Podocarpaceae en el estado de Hidalgo, México. Revista Mex. Biodivers. 77: 115118.
Contreras-Medina, R., Luna Vega, I. & Morrone, J. J. (2007). Gymnosperms and cladistic biogeography of the Mexican Transition Zone. Taxon 56: 905915.
Corral, I., Griera, A., Gómez-Gras, D., Corbella, M. & Cardellach, E. (2010). Geochemical evolution of the Panamanian Cretaceous-Paleogene volcanic arc: Insights from the Azuero Peninsula, SW Panama. Geogaceta 49: 7982.
Cuevas-Guzmán, R., Cisneros-Lepe, E. A., Jardel-Peláez, E. J., Sánchez-Rodríguez, E. V., Guzmán-Hernández, L., Núñez-López, N. M. & Rodríguez-Guerrero, C. (2011). Análisis estructural y de diversidad en los bosques de Abies de Jalisco, México. Revista Mex. Biodiv. 82: 12191233.
de Laubenfels, D. J. (1978). The genus Prumnopitys (Podocarpaceae) in Malesia. Blumea 24(1): 189190.
de Laubenfels, D. J. (1982). Podocarpaceae. In: Luces de Febres, Z. & Steyermark, J. A. (eds) Flora de Venezuela 11(2): 741. Caracas: Instituto Nacional de Parques, Dirección de Investigaciones Biológicas, Ediciones Fundación Educación Ambiental.
de Laubenfels, D. J. (1985). A taxonomic revision of the genus Podocarpus. Blumea 30: 251278.
de Laubenfels, D. J. (1991 [‘1990’]). The Podocarpaceae of Costa Rica. Brenesia 33: 119121.
de Laubenfels, D. J. (1994). Las Podocaráceas [sic] del Perú. Bol. Lima 16 (91–96): 3539.
de Laubenfels, D. J. (2007). [Curriculum vitae.], accessed 18 June 2012.
Defant, M. J., Clark, L. F., Stewart, R. H., Drummond, M. S., de Boer, J. Z., Maury, R. C., Bellon, H., Jackson, T. E. & Restrepo, J. F. (1991). Andesite and dacite genesis via contrasting processes: the geology and geochemistry of El Valle Volcano, Panama. Contrib. Mineral. Petrol. 106: 309324.
Del Castillo, R. F., Pérez de la Rosa, J. A., Vargas, G. A. & Rivera García, R. (2004). Coníferas. In: García-Mendoza, A. J., Ordóñez, M. J. & Briones-Salas, M. (eds) Biodiversidad de Oaxaca, pp. 141158. Ciudad Universitaria, México City: Instituto de Biología, UNAM. Fondo Oaxaqueño para la Conservación de la Naturaleza-World Wildlife Fund, México.
Denyer, P. & Gazel, E. (2009). The Costa Rican Jurassic to Miocene oceanic complexes: Origin, tectonics and relations. J. S. Amer. Earth Sci. 28: 429442.
Denyer, P., Baumgartner, P. O. & Gazel, E. (2006). Characterization and tectonic implications of Mesozoic-Cenozoic oceanic assemblages of Costa Rica and Western Panama. Geologica Acta 4: 219235.
Dinerstein, E., Olson, D. M., Graham, D. J., Webster, A. L., Primm, S. A., Bookbinder, M. P. & Ledec, G. (1995). A Conservation Assessment of the Terrestrial Ecoregions of Latin America and the Caribbean. Washington, DC: The World Bank.
Directorie Cartográfico de España (1998–2014). [Gazetteers and maps of Costa Rica, El Salvador, Guatemala, Honduras, México, Nicaragua and Panamá.], accessed 28 March 2014 etc.
Durand, T., Pittier, H. & Müller, J. (1891). Primitiae florae costaricensis, premier fascicule. Bull. Soc. Roy. Bot. Belg. 30: 797.
Eckenwalder, J. E. (2009). Conifers of the World: The Complete Reference. Portland, OR: Timber Press.
Enquist, C. A. F. (2002). Predicted regional impacts of climate change on the geographical distribution and diversity of tropical forests in Costa Rica. J. Biogeogr. 29: 519534.
Espindola, J. M., Zamora-Camacho, A., Godinez, M. L., Schaaf, P. & Rodríguez, S. R. (2010). The 1793 eruption of San Martín Tuxtla volcano, Veracruz, Mexico. J. Volcanol. Geotherm. Res. 197: 188208.
ESRI (Environmental Systems Resource Institute) (2008). ArcGIS 9.3. Redlands, CA: ESRI.
Estrada Chavarría, A., Rodríguez González, A. & Sánchez González, J. (2005). Evaluación y Categorización del Estado de Conservación de Plantas en Costa Rica. [Costa Rica, place of publication not stated; ?San José] Instituto Nacional de Biodiversidad, Museo Nacional de Costa Rica/SINAC.
Fa, J. E. & Morales, L. M. (1991). Mammals and protected areas in the Trans-Mexican Neovolcanic Belt. In: Mares, M. A. & Schmidly, D. J. (eds) Latin American Mammalogy: History, Biodiversity and Conservation, pp. 199226. Norman: University of Oklahoma Press.
Farjon, A. (1998). World Checklist and Bibliography of Conifers. Ed. 1. Kew: Royal Botanic Gardens. 298 pp.
Farjon, A. (2001). World Checklist and Bibliography of Conifers. Ed. 2. Kew: Royal Botanic Gardens. 309 pp.
Farjon, A. (2009). 37. Podocarpaceae. [Draft account for Flora Mesoamericana.], downloaded 28 June 2012.
Farjon, A. (2010). A Handbook of the World’s Conifers. Leiden & Boston: E. J. Brill.
Farjon, A. (2013a). Podocarpus costaricensis. In: IUCN Red List of Threatened Species, Version 2013.1., downloaded 30 August 2013.
Farjon, A. (2013b). Podocarpus matudae. In: IUCN Red List of Threatened Species, Version 2013.1., downloaded 30 August 2013.
Farjon, A. & Page, C. N. (1999). Conifers: Status Survey and Conservation Action Plan. Gland, Switzerland & Cambridge, UK: IUCN/SSC Conifer Specialist Group, IUCN.
Farris, D. W., Jaramillo, C., Bayona, G., Restrepo-Moreno, S. A., Montes, C., Cardona, A., Mora, A., Speakman, R. J., Glascock, M. D. & Valencia, V. (2011). Fracturing of the Panamanian Isthmus during initial collision with South America. Geology 39: 10071010.
Ferrari, L., López-Martínez, M., Aguirre-Díaz, G. & Carrasco-Nuñez, G. (1999). Space-time patterns of Cenozoic arc volcanism in central Mexico: From the Sierra Madre Occidental to the Mexican Volcanic Belt. Geology 27: 303306.
Flora Mesoamericana (no date). Flora Mesoamericana. Información general., accessed 1 April 2014.
Forster, R., Albrecht, H., Belisle, M., Caballero, A., Galletti, H., Lacayo, O., Ortiz, S. & Robinson, D. (2002). Comunidades forestales y mercadeo de maderas tropicales poco utilizadas de Mesoamérica. México, D.F.: Editorial Ducere, S.A., downloaded 20 February 2013.
Gaona-García, G., Sánchez-Ramos, G. & Villalón, M. L. (2009). Diagnóstico fitosanitario de la Reserva El Cielo, Tamaulipas, México. In: Cancino, E. R. & Coronado Blanco, J. M. (eds) Taller Internacional de Recursos Naturales, Memoria, 21 al 23 de Octubre del 2009, pp. 3439. Ciudad Victoria: Universidad Autónoma de Tamaulipas, UAM Agronomía y Ciencias.
Garay-Serrano, E., Bandala, V. M. & Montoya, L. (2012). Morphological and molecular identification of the ectomycorrhizal association of Lactarius fumosibrunneus and Fagus grandifolia var. mexicana trees in eastern Mexico. Mycorrhiza 22: 583588.
García Franco, J. G., Castillo-Campos, G., Mehltreter, K., Martínez, M. L. & Vázquez, G. (2008). Composición florística de un bosque mesófilo del centro de Veracruz, México. Bol. Soc. Bot. Méx. 83: 3752.
Gardner, M. (2013). Podocarpus oleifolius. In: IUCN Red List of Threatened Species, Version 2013.1., downloaded 30 August 2013.
Gaussen, H. (1976). Les Gymnospermes actuelles et fossiles. Fascicule XIV. Genre Podocarpus. Conclusion des Podocarpines. Trav. Lab. Forest. Toulouse tome 2, vol. 1, fasc. XIV. Toulouse: Faculté des Sciences.
GBIF (Global Biodiversity Information Facility) (2013). Biodiversity occurrence data (‘Podocarpus matudae: Honduras’) published by Missouri Botanical Garden. Accessed through GBIF Data Portal,, 29 April 2013.
GeoNames., accessed 31 March 2014 etc.
Gómez Pompa, A. (1964). A new Diospyros from the Misantla region in Mexico. J. Arnold Arbor. 45: 464470.
González, F.San, M. & Lavin, P. A. (1997). Los ascomycetes Acanthonitschkia, Corynelia, Lopadostoma y Camarops en México. Acta Bot. Mex. 41: 3141.
González-Espinosa, M., Ramírez-Marcial, N. & Galindo-Jaimes, L. (2006). Secondary succession in montane pine-oak forests of Chiapas, Mexico. In: Kappelle, M. (ed.) Ecology and Conservation of Neotropical Montane Oak Forests, pp. 209221. Ecological Studies, Vol. 185. Berlin & Heidelberg: Springer-Verlag.
González-Espinosa, M., Meave, J. A., Lorea-Hernández, F. G., Ibarra-Manríquez, & Newton, A. C. (2011). The Red List of Mexican Cloud Forest Trees. Cambridge, UK: Fauna and Flora International.
González-García, F. (1995). Reproductive biology and vocalizations of the Horned Guan Oreophasis derbianus in Mexico. Condor 97: 415426.
González Vace, L. A. (2011). Flora de Costa Rica I; Guía práctica. San José: Editorial Universidad Estatal a Distancia.
González-Villarreal, L. M. (2003a). Quercus tuitensis (Fagaceae, Quercus Sect. Lobatae), a new deciduous oak from western Jalisco, Mexico. Brittonia 55: 4248.
González-Villarreal, L. M. (2003b). Two new species of oak (Fagaceae, Quercus sect. Lobatae) from the Sierra Madre del Sur, Mexico. Brittonia 55: 4960.
González-Villarreal, L. M. (2006 [‘2005’]). Novelties in Clethra (Clethraceae) from Mexico. Ibugana 13(1): 1125.
Google Maps. [Co-ordinates 9°38′10″N 84°05′25″W].,-84.0902778,15z/data=!5m1!1e4, accessed 1 April 2014.
Graham, A. (1976). Studies in Neotropical paleobotany. II. The Miocene communities of Veracruz, Mexico. Ann. Missouri Bot. Gard. 63: 787842.
Graham, A. (1987). Tropical American Tertiary floras and paleoenvironments: Mexico, Costa Rica, and Panama. Amer. J. Bot. 74: 15191531.
Graham, A. (1988a). Miocene communities and paleoenvironments of southern Costa Rica. Amer. J. Bot. 74: 15011518.
Graham, A. (1988b). Studies in Neotropical paleobotany. XI. Late Tertiary vegetation and environments of southeastern Guatemala: palynofloras from the Mio-Pliocene Padre Miguel Group and the Pliocene Herrería Formation. Amer. J. Bot. 85: 14091425.
Graham, A. (1991). Studies in Neotropical paleobotany. VIII. The Pliocene communities of Panama – Introduction and ferns, gymnosperms, angiosperms (monocots). Ann. Missouri Bot. Gard. 78: 190200.
Graham, A. (1999). Studies in Neotropical paleobotany. XIII. An Oligo-Miocene palynoflora from Simojovel (Chiapas, Mexico). Amer. J. Bot. 86: 1731.
Graham, A. (2003). In the beginning: early events in the development of Mesoamerica and the lowland Maya area. In: Gómez Pompa, A., Allen, M. F., Fedick, S. L. & Jiménez-Osornio, J. J. (eds) The Lowland Maya Area: Three Millennia at the Human–Wildland Interface, pp. 3144. Binghampton, NY: Food Products Press, The Haworth Press.
Grashoff, J. L. (1974). Novelties in Stevia (Compositae: Eupatorieae). Brittonia 26: 347384.
Grayum, M. H. (1995–2013). Gazetteer of Costa Rican plant-collecting locales. St. Louis, MO: Missouri Botanical Garden., accessed 12 June 2013 and other subsequent dates.
Greuter, W., Burdet, H. M., Chaloner, W. G., Demoulin, V., Grolle, R., Hawksworth, D. L., Nicolson, D. H., Silva, P. C., Stafleu, F. A., Voss, E. G. & McNeill, J. (eds) (1988). International Code of Botanical Nomenclature (Berlin Code) adopted by the Fourteenth International Botanical Congress, Berlin, July-August 1987. Regnum Veg. 118. Königstein: Koeltz Scientific Books.
Guerrero, A. I. S. & Equihua, M. (2005). Experimental tree assemblages on the ecological rehabilitation of a cloud forest in Veracruz, Mexico. Forest Ecol. Managem. 218: 329341.
Gutiérrez, R. C. J. (2008). Proyecto CAFNET: Conectando y sosteniendo los servicios ambientales para café agroforestal, Nicaragua. Caracterización Zona Piloto Peñas Blanca. Managua: CATIE_CIRAD.
Gutiérrez-Rodríguez, C., Ornelas, J. F. & Rodríguez-Gómez, F. (2011). Chloroplast DNA phylogeography of a distylous shrub (Palicourea padifolia, Rubiaceae) reveals past fragmentation and demographic expansion in Mexican cloud forests. Molec. Phylogen. Evol. 61: 603615.
Haber, W. A. (2000). Plants and vegetation. In: Nadkarni, N. & Wheelwright, N. (eds) Monteverde, Ecology and Conservation of a Tropical Cloud Forest, pp. 3994. New York: Oxford University Press.
Halffter, G. (1987). Biogeography of the montane entomofauna of Mexico and Central America. Annu. Rev. Entomol. 32: 95114.
Heiner, A. M., Armando, E. C., Quírico, J. M. & Fabiana, M. R. (eds) (2001) Manual de Plantas Registradas en la Cuenca Hidrográfica del Río Savegre, Costa Rica. San José: INBio / Departamento de Historia Natural, Museo Nacional de Costa Rica.
Hernandez, X., E., Crum, H., Fox, W. B. & Sharp, A. J. (1951). A unique vegetational area in Tamaulipas. Bull. Torrey Bot. Club 78: 458463.
Holder, C. D. (2003). Fog precipitation in the Sierra de las Minas Biosphere Reserve, Guatemala. Hydrol. Process. 17: 20012010.
Holder, C. D. (2004). Rainfall interception and fog precipitation in a tropical montane cloud forest of Guatemala. Forest Ecol. Managem. 190: 373384.
Holwerda, F., Bruijnzeel, L. A., Muñoz-Villers, L. E., Equihua, M. & Asbjornsen, H. (2010). Rainfall and cloud water interception in mature and secondary lower montane cloud forests of central Veracruz, Mexico. J. Hydrol. 384: 8496.
Hou, C-L. & Piepenbring, M. (2009). New species and new records of Rhytismatales from Panama. Mycologia 101: 565572.
IABIN (Red Interamericana de Información sobre Biodiversidad)., accessed 28 June 2012.
IPNI (The International Plant Names Index)., accessed 1 April 2014 etc.
Iremonger, S., Liesner, R. & Sayre, R. (1995). Plant records from natural forest communities in the Bladen Nature Reserve, Maya Mountains, Belize. Caribbean J. Sci. 31: 3048.
Ishiki, M. (1988). Las selvas bajas perennifolias del Cerro Salomón, región de Chimalapa, Oaxaca. Flora, comunidades y relaciones fitogeográficas. Tesis de maestría, Colegio de Posgraduados, Chapingo.
IUCN (2012). IUCN Red List Categories and Criteria: Version 3.1. Ed. 2. Gland, Switzerland and Cambridge, UK: IUCN.
IUCN Standards and Petitions Subcommittee (2013). Guidelines for using the IUCN Red List Categories and Criteria. Version 10.1 (September 2013)., downloaded 19 November 2013.
James, K. (2006). Arguments for and against the Pacific origin of the Caribbean Plate: Discussion, finding for an inter-American origin. Geol. Acta 4: 279302.
James, K. H. (2009a). In situ origin of the Caribbean: discussion of data. Geol. Soc., London, Spec. Publ. 328: 77125.
James, K. H. (2009b). Evolution of Middle America and the in situ Caribbean Plate model. Geol. Soc., London, Spec. Publ. 328: 127138.
Janzen, D. H. (1967). Synchronization of sexual reproduction of trees within the dry season in Central America. Evolution 21: 620637.
Jiménez, Q. (1999a). Podocarpus costaricensis., accessed 20 February 2013.
Jiménez, Q. (1999b). Podocarpus guatemalensis., accessed 20 February 2013.
Kappelle, M. (2006). Structure and composition of Costa Rican montane oak forests. In: Kappelle, M. (ed.) Ecology and Conservation of Neotropical Montane Oak Forests, pp. 127139. Ecological Studies Vol. 185. Berlin & Heidelberg: Springer-Verlag.
Kappelle, M., Van Uffelen, J-G. & Cleef, A. M. (1995). Altitudinal zonation of montane Quercus forests along two transects in Chirripó National Park, Costa Rica. Vegetatio 119: 119153.
Kappelle, M., Avertin, G., Juárez, M. E. & Zamora, N. (2000). Useful plants within a campesino community in a Costa Rican montane cloud forest. Mountain Res. Developm. 20: 162171.
Karsten, G. (1897). Notizen über einige mexikanische Pflanzen. Ber. Deutsch. Bot. Ges. 15: 1016.
Kellman, M., Tackaberry, R. & Rigg, L. (1998). Structure and function in two tropical gallery forest communities: implications for forest conservation in fragmented systems. J. Appl. Ecol. 35: 195206.
Keppie, D. F. (2012). Derivation of the Chortis and Chiapas blocks from the western Gulf of Mexico in the latest Cretaceous–Cenozoic: the Pirate model. Int. Geol. Rev. 54: 17651775.
Keppie, D. F. & Keppie, J. D. (2012). An alternative Pangea reconstruction for Middle America with the Chortis Block in the Gulf of Mexico: tectonic implications. Int. Geol. Rev. 54: 16851696.
Keppie, J. D. & Morán-Zenteno, D. J. (2005). Tectonic implications of alternative Cenozoic reconstructions for southern Mexico and the Chortis Block. Int. Geol. Rev. 47: 473491.
Kerhoulas, N. J. & Arbogast, B. S. (2010). Molecular systematics and Pleistocene biogeography of Mesoamerican flying squirrels. J. Mammalogy 91: 654667.
Kim, YH., Clayton, R. W. & Keppie, F. (2011). Evidence of a collision between the Yucatán Block and Mexico in the Miocene. Geophys. J. Int. 187: 9891000.
Knopf, P., Schulz, C., Little, D. P., Stützel, T. & Stevenson, D. W. (2012). Relationships within Podocarpaceae based on DNA sequence, anatomical, morphological, and biogeographical data. Cladistics 28: 271299.
Kuijt, J. & Kellogg, E. A. (1996). Miscellaneous mistletoe notes, 20–36. Novon 6: 3353.
Lachniet, M. S. & Roy, A. R. (2011). Chapter 60. Costa Rica and Guatemala. Dev. Quat. Sci. 15: 843848.
Lambert, A. B. (1824). A description of the genus Pinus. Ed. 1, vol. 2 [by D. Don]. London: J. White.
Linares, J. L. (2005 [‘2003’]). Listado comentado de los árboles nativos y cultivados en la república de El Salvador. Ceiba 44(2): 105268.
Llorente-Bousquets, J. & Escalante-Pliego, P. (1992). Insular biogeography of submontane humid forests in Mexico. In: Darwin, S. P. & Welden, A. L. (eds) Biogeography of Mesoamerica, pp. 139146. Tulane Univ. Stud. Zool. Bot., Suppl. Publ. 1.
Luna Vega, I., Alcántara-Ayala, O., Contreras-Medina, R. & Ponce Vargas, A. (2006). Biogeography, current knowledge and conservation of threatened vascular plants characteristic of Mexican temperate forests. Biodivers. Conserv. 15: 37733799.
Lundell, C. L. (1937). Studies of Mexican and Central American plants – II(a). Phytologia 1: 212222.
Macey, A. (1975). The vegetation of Volcán Poás National Park, Costa Rica. Revista Biol. Trop. 23: 239255., downloaded 20 June 2012.
Magaña, V., Amador, J. A. & Medina, S. J. (1999). The midsummer drought over Mexico and Central America. J. Climate 12: 15771588.
Marshall, C. J. & Liebherr, J. K. (2000). Cladistic biogeography of the Mexican Transition Zone. J. Biogeogr. 27: 203216.
Marshall, J. S. (2007). Geomorphology and physiographic provinces of Central America. In: Bundschuh, J. & Alvarado, G. E. (eds) Central America: Geology, Resources, and Natural Hazards 1: 75122. London, etc.: Taylor & Francis.
Marshall, J. S., Idleman, B. D., Gardner, T. W. & Fisher, D. M. (2003). Landscape evolution within a retreating volcanic arc, Costa Rica, Central America. Geology 31: 419422.
Martens, U., Weber, B. & Valencia, V. A. (2010). U/Pb geochronology of Devonian and older Paleozoic beds in the southeastern Maya block, Central America: Its affinity with peri-Gondwanan terranes. Geol. Soc. Amer. Bull. 122: 815829.
Martin, P. S. (1958). A biogeography of reptiles and amphibians in the Gómez Farías Region, Tamaulipas, Mexico. Misc. Publ. Mus. Zool. Univ. Michigan, No. 101. Ann Arbor: University of Michigan.
Martín, R. A. (1993). Podocarpaceae. In: Killeen, T. J., Emilia García, E. & Beck, S. G. (eds) Guía de arboles de Bolivia, pp. 641645. La Paz: Herbario Nacional de Bolivia, & St. Louis: Missouri Botanical Garden.
Martínez-Camilo, R., Martínez-Melendez, N., Perez-Farrera, M. A. & Lorence, D. H. (2011). Una nueva especie de Pseudomiltemia (Rubiaceae) de Chiapas, México. Brittonia 63: 197202.
Martínez-Meléndez, J., Pérez-Farrera, M. Á. & Farrera-Sarmiento, O. (2008). Inventario florístico del Cerro El Cebú y zonas adyacentes en la Reserva de la Biosfera El Triunfo (Polígono V), Chiapas, México. Bol. Soc. Bot. Méx. 82: 2140.;script=sci_arttext, downloaded 4 October 2013.
Matamoros, Y. & Seal, U. S. (eds) (1996). Report of Threatened Plants of Costa Rica Workshop, 4–6 October. Apple Valley, MN: IUCN/SSC Conservation Breeding Specialist Group.
Matson, J. O., Ordóñez-Garza, N., Bulmer, W. & Eckerlin, R. P. (2012). Small mammal communities in the Sierra de los Cuchumatanes, Huehuetenango, Guatemala. Mastozool. Neotrop. 19: 7184.
Matuda, E. (1950a). A contribution to our knowledge of wild flora of Mt. Ovando. Amer. Midl. Nat. 43: 195223.
Matuda, E. (1950b). A contribution to our knowledge of the wild and cultivated flora of Chiapas – 1. Districts Soconusco and Mariscal. Amer. Midl. Nat. 44: 513616.
McNeill, J., Barrie, F. R., Buck, W. R., Demoulin, V., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Marhold, K., Prado, J., Prud’homme van Reine, W. F., Smith, G. F., Wiersema, J. H. & Turland, N. J. (2012). International Code of Nomenclature for algae, fungi, and plants (Melbourne Code) adopted by the Eighteenth International Botanical Congress Melbourne, Australia, July 2011. Regnum Veg. 154. Ruggell: A. R. G. Gantner Verlag KG.
McVaugh, R. (1966). The occurrence of the genus Podocarpus in western Mexico. Sobretiro Ci. Mex. 24(5–6): 223226.
McVaugh, R. & Schmid, R. (1967). Novelties in Satureia Sect. Gardoquia (Labiatae). Brittonia 19: 261267.
Mejía, V., D. A. (2001). Honduras. In: Kappelle, M. & Brown, A. D. (eds) Bosques Nublados del Neotrópico, pp. 243282. Santo Domingo de Heredia: Instituto Nacional de Biodiversidad.
Merello, M. (2003). Podocarpaceae. In: Hammel, B. E., Grayum, M. H., Herrera, C. & Zamora, N. (eds) Manuel de Plantas de Costa Rica. Volumen II. Gimnospermas y Monocotiledóneas (Agavaceae–Musaceae), pp. 1113. St. Louis, MO: Missouri Botanical Garden Press.
Merker, C. A., Barbour, W. R., Scholten, J. A. & Dayton, W. A. (1943). The Forests of Costa Rica. Washington: USDA Forest Service.
Mill, R. R. (2015). A revision of the genus Podocarpus (Podocarpaceae): II. The species of the Caribbean Bioregion. Edinburgh J. Bot. 72: 61185.
Molina, R., A. (1964). Coniferas de Honduras. Ceiba 10(1): 521.
Monterroso-Rivas, A. I., Gómez-Díaz, J. D. & Tinoco-Rueda, J. A. (2013). Bosque mesófilo de montaña y escenarios de cambio climático: una evaluación en Hidalgo, México. Revista Chapingo, Ser. Ci. Forest. Ambiente 19(1): 2943.
Monterroso Salinas, J. F. (1976). Analisis floristico y estructural del biotopo protegido de Purulha para la conservación del Quetzal. San Carlos: Thesis, Facultad de Ciencias Quimícas y Farmacia, Universidad de San Carlos.
Montes, C., Cardona, A., McFadden, R., Morón, S. E., Silva, C. A., Restrepo-Moreno, S., Ramírez, D. A., Hoyos, N., Wilson, J., Farris, D., Bayona, G. A., Jaramillo, C. A., Valencia, V., Bryan, J. & Flores, J. A. (2012). Evidence for middle Eocene and younger land emergence in central Panama: Implications for Isthmus closure. Geol. Soc. Amer. Bull. 124: 780799.
Morán-Zenteno, D. J., Cerca, M. & Keppie, J. D. (2007). The Cenozoic tectonic and magmatic evolution of southwestern México: Advances and problems of interpretation. In: Alaniz-Álvarez, S. A. & Nieto-Samaniego, Á. F. (eds) Geology of México: Celebrating the Centenary of the Geological Society of México. Geol. Soc. Amer. Spec. Pap. 422: 7191.
Morrone, J. J. (2001a). Toward a cladistic model for the Caribbean Subregion: delimitation of areas of endemism. Caldasia 23: 4376.
Morrone, J. J. (2001b). Biogeografia de América Latina y el Caribe. M&T – Manuales y Tesis SEA Vol. 3. Zaragoza: Sociedad Entomológica Aragonesa (SEA).
Morrone, J. J. (2006). Biogeographic areas and transition zones of Latin America and the Caribbean Islands based on panbiogeographic and cladistic analyses of entomofauna. Annu. Rev. Entomol. 51: 467494.
Morrone, J. J., Espinosa Organista, D. & Llorente Bousquets, J. (2002). Mexican biogeographic provinces: preliminary scheme, general characterizations, and synonymies. Acta Zool. Mex. (n.s.) 85: 83108.
Moya, R., Wiemann, M. C. & Olivares, C. (2013). Identification of endangered or threatened Costa Rican tree species by wood anatomy and fluorescence activity. Rev. Biol. Trop. (Int. J. Trop. Biol.) 61: 11331156.
Muñoz-Villers, L. E. & López-Blanco, J. (2007). Land use/cover changes using Landsat TM/ETM images in a tropical and biodiverse mountainous area of central-eastern Mexico. Int. J. Remote Sensing 29: 7193.
Negrete-Yankelevich, S., Fragoso, C., Newton, A. C. & Heal, O. W. (2007). Successional changes in soil, litter and macroinvertebrate parameters following selective logging in a Mexican Cloud Forest. Appl. Soil Ecol. 35: 340355.
Nelson, S. A. & Gonzalez-Caver, E. (1992). Geology and K-Ar dating of the Tuxtla Volcanic Field, Veracruz, Mexico. Bull. Volcanol. 55: 8596.
Newstrom, L. E., Frankie, G. W. & Baker, H. G. (1994). A new classification for plant phenology based on flowering patterns in lowland tropical rain forest trees at La Selva, Costa Rica. Biotropica 26: 141159.
Niembro Rocas, A. (1986). Arboles y arbustos útiles de México. México, D.F.: LIMUSA.
Oldfield, S., Lusty, C. & MacKinven, A. (1998). The World List of Threatened Trees. Cambridge, UK: WCMC/World Conservation Press.
Opler, P. A., Frankie, G. W. & Baker, H. G. (1980). Comparative phenological studies of treelet and shrub species in tropical wet and dry forests in the lowlands of Costa Rica. J. Ecol. 68: 167188.
Ornelas, J. F., Ruiz-Sánchez, E. & Sosa, V. (2010). Phylogeography of Podocarpus matudae (Podocarpaceae): pre-Quaternary relicts in northern Mesoamerican cloud forests. J. Biogeogr. 37: 23842396.
Ornelas, J. F., Sosa, V., Soltis, D. E., Daza, J. M., González, C., Soltis, P. S., Gutiérrez-Rodríguez, C., Espinosa de los Monteros, A., Castoe, T. A., Bell, C. & Ruiz-Sanchez, E. (2013). Comparative phylogeographic analyses illustrate the complex evolutionary history of threatened cloud forests of northern Mesoamerica. PLoS ONE 8(2): e568283. doi:10.1371/journal.pone.0056283, downloaded 8 October 2013.
Ortega-Gutiérrez, F., Solari, L. A., Ortega-Obregón, C., Elías-Herrera, M., Martens, U., Morán-Icál, S., Chiquín, M., Keppie, J. D., Torres De León, R. & Schaaf, P. (2007). The Maya-Chortís boundary: a tectonostratigraphic approach. Int. Geol. Rev. 49: 9961024.
Padilla y Sánchez, R. J. (2007). Evolución geológica del sureste mexicano desde el Mesozoico al presente en el contexto regional del Golfo de México. Bol. Soc. Geol. Mex. 59: 1942.
Parker, T., Carrión, J. & Samudio, R. (2004). Section 118/119. Biodiversity and Tropical Forestry Assessment of the USAID/Panama Program. Environment, Biodiversity, Water, and Tropical Forest Conservation, Protection, and Management in Panama: Assessment and Recommendations. BIOFOR IQC No. LAG-I-00-99-00014-00, Task Order #824. [Washington, DC:] Chemonics International.
Parlatore, P. (1868). Podocarpus. In: De Candolle, A., Prodromus systematis vegetabilium 16(2): 507521.
Pedraza, R. A. & Williams-Linera, G. (2003). Evaluation of native tree species for the rehabilitation of deforested areas in a Mexican cloud forest. New Forests 26: 8399.
Penn, M. G., Sutton, D. A. & Monro, A. (2004). Vegetation of the Greater Maya Mountains, Belize. Syst. Biodiv. 2: 2144.
Pérez de la Rosa, J. (1999). Regional Action Plan: Conifers threatened in Nueva Galicia, Mexico. In: Farjon, A. & Page, C. N., Conifers: Status Survey and Conservation Action Plan, pp. 5558. Gland, Switzerland & Cambridge, UK: IUCN/SSC Conifer Specialist Group, IUCN.
Pérez Ribera, R. (2002). Evaluación del impacto del aprovechamiento Forestal en Los Mogos, Península de Osa, Costa Rica. Cartago: Instituto Tecnológico de Costa Rica., downloaded 20 February 2013.
Pilger, R. (1903). Taxaceae. Being Engler, A. (ed.) Das Pflanzenreich IV.5 (Heft 18). Leipzig: Wilhelm Engelmann.
Pineda, A. G. (2006). Flora Util. Etnobotánica de Nicaragua. Managua: Ministerio del Ambiente y los Recursos Naturales – MARENA., downloaded 20 February 2013.
Portig, W. H. (1965). Central American rainfall. Geogr. Rev. 55: 6890.
Puig, H., Bracho, R. & Sosa, V. (1983). Composición floristica y estructura del bosque mesofilo en Gómez Farías, Tamaulipas, Mexico. Biotica 8(4): 339359.
Quesada Monge, R. (2004). Especies forestales vedadas y bajo otras categorías de protección en Costa Rica. Kurú: Revista Forestal (Costa Rica) 1(2): 15., downloaded 6 February 2012.
Quesada Monge, R. & Fernández Vega, J. (2005a). Actualización de listado de especies arbóreas de uso forestal y otros usos en Costa Rica. Listado por nombres científicos. Kurú: Revista Forestal (Costa Rica) 2(4): 145.
Quesada Monge, R. & Fernández Vega, J. (2005b). Actualización de listado de especies arbóreas de uso forestal y otros usos en Costa Rica. Listado por nombres comunes. Kurú: Revista Forestal (Costa Rica) 2(5): 144.
Ramos Álvarez, C. H., Martínez, E., Nava-Cruz, Y., Martínez-Bravo, R. & Ricker, M. (2008). Descripción de la vegetación y diagnóstico de la diversidad florística en el Parque Ecológico Jaguaroundi. In: Nava, Y. & Rosas, I. (eds) El Parque Ecologico Jaguaroundi: Conservacion de la selva tropical veracruzana en una zona industrializada, pp. 5578. Ciudad Universitaria, México City: Programa Universitario de Medio Ambiente, Universidad Nacional Autonoma de México.
Rasband, W. S. (1997–2014). ImageJ. Bethesda, MD: U.S. National Institutes of Health., accessed 27 August 2011 onwards.
Rauscher, S. A., Giorgi, F., Diffenbaugh, N. S. & Seth, A. (2008). Extension and intensification of the Meso-American mid-summer drought in the twenty-first century. Clim. Dyn. 31: 551571.
Reeves, R. D., Baker, A. J. & Romero, M. R. (2007). The ultramafic flora of the Santa Elena peninsula, Costa Rica: A biogeochemical reconnaissance. J. Geochem. Explor. 93: 153159.
Reiche, K. F. (1927). Podocarpus guatemalensis, el Palmillo. México Forest. 5: 7778.
Restrepo, C. & Alvarez, N. (2006). Landslides and their contribution to land-cover change in the mountains of Mexico and Central America. Biotropica 38: 446457.
Reyes-Castillo, P. (2000). Coleoptera Passalidae de México. In: Martín-Piera, F., Morrone, J. J. & Melic, A. (eds) m3m: Monografías Tercer Milenio 1: 171182. Zaragoza: SEA.
Rodríguez, S. R., Morales-Barrera, W., Layer, P. & González-Mercado, E. (2010). A Quaternary monogenetic volcanic field in the Xalapa region, eastern Trans-Mexican volcanic belt: geology, distribution and morphology of the volcanic vents. J. Volcanol. Geotherm. Res. 197: 149166.
Rodriguez Martinez, A. (1999). Condiciones ambientales que favorecen la distribución natural de Podocarpus guatemalensis Standley en el litoral Atlántico de Honduras. Siguatepeque: Ing. For. thesis, Escuela Nacional de Ciencias Forestales. Abstract available at, accessed 20 February 2013.
Rogers, R. D. (1998). Geology along the Río Patuca and Río Wampú Honduras., accessed 5 November 2013.
Rogers, R. D., Mann, P. & Emmet, P. A. (2007). Tectonic terranes of the Chortis block based on integration of regional aeromagnetic and geologic data. Geol. Soc. Amer. Spec. Pap. 428: 6588.
Rojas-Soto, O. R., Sosa, V. & Ornelas, J. F. (2012). Forecasting cloud forest in eastern and southern Mexico: conservation insights under future climate change scenarios. Biodivers. Conserv. 21: 26712690.
Romero, C. (1999). Reduced-impact logging effects on commercial non-vascular pendant epiphyte biomass in a tropical montane forest in Costa Rica. Forest Ecol. Managem. 118: 117125.
Romero Velásquez, T. L. (2009). Cartografía y caracterización de ecosistemas de la Subcuenca del Río Cumes, Jesús de Otoro, Intibucá, Honduras. Proyecto especial del programa de Ingeniería en Desarrollo Socioeconómico y Ambiente, Zamorano, Honduras., accessed 7 October 2013.
Rosenfeld, J. H. (1990). Sedimentary rocks of the Santa Cruz ophiolite, Guatemala – a proto-Caribbean history. Trans. 12th Caribbean Geol. Conf., pp. 513519. South Miami: Miami Geological Society.
Ruiz, C. (2006). Plan Maestro de la Reserva de Manantiales Cerro San Gil. Guatemala: Consejo Nacional de Áreas Protegidas, Fundación para el Ecodesarrollo y la Conservación & The Nature Conservancy.
Rzedowski, J. & McVaugh, R. (1966). La vegetación de Nueva Galicia. Contr. Univ. Michigan Herb. 9: 1123.
Sánchez-Ramos, G. (1999). Herbivory in Podocarpus reichei Buchholz et Gray (Podocarpaceae) by Anisodes gigantula (Lepidoptera: Geometridae) in mesophilic forest at Gomez Farias, Tamaulipas, Mexico. / Herbivoría en Podocarpus reichei Buchholz et Gray (Podocarpaceae) por Anisodes gigantula (Lepidoptera: Geometridae) en el bosque mesofilo de Gómez Farias, Tamaulipas, México. Brenesia 51: 45–57.
Sánchez-Ramos, G., Dirzo, R. & Balcázar-Lara, M. A. (1999). Especifidad y herbivoría de Lepidoptera sobre especies pioneras y tolerantes del bosque mesofilo de la Reserva de la Biosfera El Cielo, Tamaulipas, Mexico. Acta Zool. Mex. (n.s.) 78: 103118.
San Emeterio, L. & Cavero, R. Y. (2000). Contribución a la flora de las cuencas Juan de Paz y Las Cañas, Reserva de Biosfera Sierra de las Minas (Guatemala). Publ. Biol. Univ. Navarra, Ser. Bot. 13: 85100.
Sharp, A. J. (1946). La distribución del género Podocarpus en México (1). Bol. Soc. Bot. México 4: 1718.
Sharp, A. J. (1949). Distribución del género Podocarpus en México. Bol. Soc. Bot. México 8: 25.
Silba, A. D. & de Silva, J. A. (2014). Yon nouvo nan Podocarpus oleifolius D. Don (Podocarpaceae) soti nan Columbia. J. Int. Conifer Preserv. Soc. 21(1): 1930.
Silba, J. (1984). An international census of the Coniferae, I. Phytologia Mem. 7. Plainfield, NJ: Harold N. Moldenke & Alma L. Moldenke.
Silba, J. (1986). Encyclopaedia Coniferae. Phytologia Mem. 8. Corvallis, OR: Harold N. Moldenke & Alma L. Moldenke.
Silba, J. (1990). A supplement to the International Census of the Coniferae, II. Phytologia 68: 778; corrections, Phytologia 68: 512.
Silba, J. (2000). Variation geographic et populations isole de les gymnospermes rarissime. J. Int. Conifer Preserv. Soc. 7(1): 1740.
Silba, J. (2010). An international census of the Coniferae. Part II. Podocarpus. J. Int. Conif. Preserv. Soc. 17(1): 719, 19 [bis], 20.
Solano Quintero, J. & Villalobos Flores, R. (2001). Aspectos fisiográficos aplicados a un bosquejo de regionalización geográfico climático de Costa Rica. Top. Meteor. Oceanog. 8: 2639.
Solórzano, S., Castillo, S., Valverde, T. & Ávila, L. (2000). Quetzal abundance in relation to fruit availability in a cloud forest in Southeastern Mexico. Biotropica 32: 523532.
Standley, P. C. (1924). Nine new species of plants from Central America. Proc. Biol. Soc. Washington 37: 4954.
Standley, P. C. (1937). Flora of Costa Rica. Part 1. Field Mus. Nat. Hist., Bot. ser. 18. Chicago: Field Museum of Natural History. [4 parts, 1937–1938; Podocarpaceae part 1, pp. 64–65, 12 Oct. 1937.]
Standley, P. C. & Steyermark, J. A. (1958). Flora of Guatemala. Podocarpaceae. Fieldiana, Bot. 24(1): 2023. Chicago: Chicago Natural History Museum. [Taxonomic account of Podocarpaceae is by N. E. Gray.]
Stark Schilling, D. (2004). Taxonomic studies of Caribbean and Central American species of Podocarpus subgenus Podocarpus: A multi-disciplinary approach. Unpublished MSc thesis, Royal Botanic Garden Edinburgh/University of Edinburgh.
Styles, B. T. & Hughes, C. E. (1983). Studies of variation in Central American pines III. Notes on the taxonomy and nomenclature of the pines and related gymnosperms in Honduras and adjacent Latin American Republics. Brenesia 21: 269291.
Takhtajan, A. (1986; transl. Crovello, T. J., ed. Cronquist, A.). Floristic Regions of the World. Berkeley: University of California Press.
Tassi, F., Vaselli, O., Capaccioni, B., Giolito, C., Duarte, E., Fernandez, E., Minissale, A. & Magro, G. (2005). The hydrothermal-volcanic system of Rincon de la Vieja volcano (Costa Rica): A combined (inorganic and organic) geochemical approach to understanding the origin of the fluid discharges and its possible application to volcanic surveillance. J. Volcanol. Geotherm. Res. 148: 315333.
Taylor, B. W. (1963). An outline of the vegetation of Nicaragua. J. Ecol. 51: 2754.
Taylor, M. A. & Alfaro, E. J. (2005). Central America and the Caribbean, Climate of. In: Oliver, J. E. (ed.) Encyclopedia of World Climatology, pp. 183189.
Thiers, B. [continuously updated]. Index Herbariorum: A global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium., accessed 31 March 2014 etc.
Thomas, P. & Farjon, A. (2013). Podocarpus guatemalensis. In: IUCN (2013). IUCN Red List of Threatened Species, Version 2013.1., downloaded 30 August 2013.
Todzia, C. A. (1999). Ten new species of Tibouchina (Melastomataceae) from Mexico. Brittonia 51: 255279.
Toledo-Aceves, T., Meave, J. A., González-Espinosa, M. & Ramírez-Marcial, N. (2011). Tropical montane cloud forests: Current threats and opportunities for their conservation and sustainable management in Mexico. J. Environm. Managem. 92: 974981.
Tonduz, A. (1896). Herborisations au Costa-Rica. III. Le bassin du Diquis. Bull. Herb. Boissier 4: 163177.
Torres-Romero, J. H. (1988). Podocarpaceae. In: Pinto, P. & Lozano, G. (eds) Flora Colombia Monogr. 5: 567. Bogotá: Instituto de Ciencias Naturales, Museo de Historia Natural, Faculdad de Ciencias, Universidad Nacional de Colombia.
Townsend, J. H., Butler, J. M., Wilson, L. D. & Austin, J. D. (2009). A new species of salamander in the Bolitoglossa dunni group (Caudata: Plethodontidae: Bolitoglossinae) from Parque Nacional Montaña de Yoro, Honduras. Salamandra 45: 95105., downloaded 20 February 2013.
TROPICOS. , accessed on various dates e.g. 27 January 2015.
Valdés, O. T., Franco, G. F., Rodríguez, A. M., Flores, R. E. G., Hernández, G. S., Rodríguez, R. R., Mariani, A. D. & Calzada, I. (1995). Listados florísticos de México. XII. Flora de la Reserva Ecológica Sierra de San Juan, Nayarit, México. Instituto de Biología, Universidad Nacional Autónoma de México.
Vannucchi, P., Fisher, D. M., Bier, S. & Gardner, T. W. (2006). From seamount accretion to tectonic erosion: Formation of Osa Mélange and the effects of Cocos Ridge subduction in southern Costa Rica. Tectonics, 25: TC2004. 19 pp. doi:10.1029/2005TC001855, downloaded 3 October 2013.
Vázquez, A. & Cuevas, R. (1989). Una nueva especie tropical de Populus (Salicaceae) de la Sierra de Manantlán, Jalisco, Mexico. Acta Bot. Mex. 8: 3945.
Vázquez-García, J. A. (1994). Magnolia (Magnoliaceae) in Mexico and Central America: a synopsis. Brittonia 46: 123.
Vázquez García, J. A., Cuevas, G., R., Cochrane, T. S., Iltis, H. H., Santana, M., F. J. &Guzman, H., L. (1995). Flora de Manantlán. Sida, Botanical Miscellany no. 13. Fort Worth: Botanical Research Institute of Texas.
Vázquez-García, J. A., Pérez-Farrera, M. Á., Martínez-Meléndez, N., Nieves-Hernández, G. & Muñiz-Castro, M. Á. (2012). Magnolia mayae (Magnoliaceae), a new species from Chiapas, Mexico. Bot. Sci. 90: 109112.
Venable, M. (1994). A geological, tectonic, and metallogenetic evaluation of the Siuna terrane (Nicaragua). Tucson, AZ: Unpublished PhD dissertation, University of Arizona., downloaded 8 November 2013.
Vicuña-Miñano, E. E. (2005). Las Podocarpáceas de los bosques montanos del noroccidente peruano. Podocarpaceae of the montane forests from northwestern Peru. Rev. Peru. Biol. 12: 283288.
Vicuña-Miñano, E. & León, J. M. (2003). Notas sobre Podocarpáceas de cuatro bosques montanos de la Provincia de San Ignacio – Cajamarca, Perú. Arnaldoa 10: 1943.
Vivero, J. L., Szejner, M., Gordon, J. & Magin, G. (2006). The Red List of Trees of Guatemala. Cambridge, UK: Fauna & Flora International.
Vogelmann, H. W. (1973). Fog precipitation in the cloud forests of eastern Mexico. BioScience 23: 96100.
Vovides, A. P., Luna, V. & Medina, G. (1997). Relacion de algunas plantas y hongos mexicanos raros, amenazados o en peligro de extincion y sugerencias para su conservación. Acta Bot. Mex. 39: 142.
Walper, J. L. (1960). Geology of Coban-Purulha Area, Alta Verapaz, Guatemala. AAPG Bull. 44: 12731315.
Whattam, S. A., Montes, C., McFadden, R. R., Cardona, A., Ramirez, D. & Valencia, V. (2012). Age and origin of earliest adakitic-like magmatism in Panama: Implications for the tectonic evolution of the Panamanian magmatic arc system. Lithos 142–143: 226244.
Wildfinder., accessed 28 March 2014 etc.
Williams, L. G. (1991). Nota sobre la estructura del estrato arbóreo del bosque mesófilo de montaña en los alrededores del campamento El Triunfo, Chiapas. Acta Bot. Mex. 13: 17.
Williams, M. D. (1975). Emplacement of Sierra de Santa Cruz eastern Guatemala. AAPG Bull. 59: 12111216.
Williams, W. L. W. (ed.) (2009). Resultados del Inventário Nacional Forestal: Nicaragua 2007–2008/INAFOR. Managua: Instituto Nacional Forestal.
Williams-Linera, G., Toledo-Garibaldi, M. & Hernández, C. G. (2013). How heterogeneous are the cloud forest communities in the mountains of central Veracruz, Mexico? Pl. Ecol. 214: 685701.
Witt, C., Rangin, C., Andreani, L., Olaez, N. & Martinez, J. (2012a). The transpressive left-lateral Sierra Madre de Chiapas and its buried front in the Tabasco plain (southern Mexico). J. Geol. Soc. 169: 143155.
Witt, C., Brichau, S. & Carter, A. (2012b). New constraints on the origin of the Sierra Madre de Chiapas (south Mexico) from sediment provenance and apatite thermochronometry. Tectonics 31: TC6001. 15 pp. doi:10.1029/2012TC003141, downloaded 1 November 2013.
Wood, S. L. (1969). New records and species of Neotropical bark beetles (Scolytidae: Coleoptera). Part IV. Sci. Bull. Brigham Young Univ., Biol. Ser. 10(2): 146.
Wood, S. L. (1974). New species of American Corthylus (Coleoptera: Scolytidae). Great Basin Naturalist 34: 181202.
Wood, S. L. (1982). The bark and ambrosia beetles of North and Central America, a taxonomic monograph. Great Basin Naturalist Memoirs no. 6.
Wood, S. L., Stevens, G. C. & Lezama, H. J. (1991). Los Scolytidae de Costa Rica: clave de géneros y de la subfamilia Hylesinae (Coleoptera). Rev. Biol. Trop. 39: 125148.
Wood, S. L., Stevens, G. C. & Lezama, H. J. (1992). Los Scolytidae (Coleoptera) de Costa Rica: Clave de la subfamilia Scolytinae, Tribu Corthylini. Rev. Biol. Trop. 40: 247286.
Woodson, R. E. & Schery, R. W. (1941). Contributions toward a flora of Panama. V. Collections chiefly by Paul H. Allen, and by Robert E. Woodson, Jr. and Robert W. Schery. Ann. Missouri Bot. Gard. 28: 409491.
Zamudio, S. (2002). Flora del Bajío y de regiones adyacentes. Fasciculo 105. Familia Podocarpaceae. Patzcuaro, Michoacán: Instituto de Ecologia., downloaded 15 February 2012.



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