This paper addresses the questions of what we have learned about how and when dense star clusters form, and what studies of star clusters have revealed about galaxy formation and evolution. One important observation is that globular clusters are observed to form in galaxy mergers and starbursts in the local universe, which both provides constraints on models of globular cluster formation, and suggests that similar physical conditions existed when most early-type galaxies and their globular clusters formed in the past. A second important observation is that globular cluster systems typically have bimodal color distributions. This was predicted by merger models, and indicates an episodic formation history for elliptical galaxies. A third and very recent result is the discovery of large populations of intermediate age globular clusters in several elliptical galaxies through the use of optical to near-infrared colors. These provide an important link between young cluster systems observed in starbursts and mergers and old cluster systems. This continuum of ages of the metal-rich globular cluster systems also indicates that there is no special age or epoch for the formation of the metal-rich globular clusters, which comprise about half of the cluster population. The paper concludes with a brief discussion of recent results on the globular cluster – low-mass X-ray binary connection.