The genus Lycaota was placed by MacGillivray (1908) in a separate subfamily closely related to the Tenthredininae. Rohwer in 1911, transferred it to the subfamily Blennocampinae (Empriinae of Rohwer) on the basis of the position of the medio-cubital cross-vein. A study of the Selandriinae, Emphytinae and Blennocampinae indicates very strongly that the three subfamilies form a compact unit and that the predicellate condition of the first anal cell in the Blennocampinae arose from the condition existing in the Selandriinae and Emphytinae, not from the condition exhibited by Lycaota, which is very typical of the condition found in a large number of the Tenthredininae.