Skip to main content Accessibility help
×
Hostname: page-component-5c6d5d7d68-wpx84 Total loading time: 0 Render date: 2024-08-18T00:06:01.360Z Has data issue: false hasContentIssue false

3 - Faunal composition and distribution of mammals in western coniferous forests

Published online by Cambridge University Press:  15 December 2009

Timothy E. Lawlor
Affiliation:
Dept. Biological Sciences, Humboldt State University, Arcata, California 95521, USA
Cynthia J. Zabel
Affiliation:
Pacific Southwest Research Station
Robert G. Anthony
Affiliation:
Oregon State University
Get access

Summary

Mammals are prominent features of the landscape in coniferous forests of western North America. Large herbivores such as deer, elk, moose, and caribou are particularly noticeable. Smaller, though no less conspicuous, mammals include diurnal ground squirrels, marmots, pikas, and numerous species of chipmunks, and nocturnal bats. Overall, the mammalian faunas of western coniferous forests are rich and varied. Knowledge of their taxonomic makeup and biogeography provides a foundation for understanding current patterns of distribution, habitat use, and management. An examination of the distribution and diversity patterns of coniferous-forest species, together with an assessment of the factors influencing these patterns, forms the basis for this review.

Faunal composition

Nearly half (194) of approximately 400 species of mammals occurring in North America north of Mexico occupy western coniferous forests and associated meadow, brush, or riparian habitats (Table 3.1, Appendix). Eight orders and 27 families of mammals are represented. Per unit area, western North America has a much higher species richness than other temperate or boreal areas of the continent. This difference is readily discernible when comparing western and eastern coniferous forests (Hallett et al. 2003). Chipmunks (Tamias spp.) constitute a striking example: 21 species of chipmunks occupy western coniferous forests; there is only one (Tamias striatus) in eastern forests. Similar, though less dramatic examples, occur among shrews (Sorex spp.), bats (Myotis spp.), ground squirrels (Spermophilus spp., Marmota spp.), and other rodents (e.g., Peromyscus spp.).

The list of species (Table 3.1, Appendix) is somewhat loosely drawn.

Type
Chapter
Information
Mammal Community Dynamics
Management and Conservation in the Coniferous Forests of Western North America
, pp. 41 - 80
Publisher: Cambridge University Press
Print publication year: 2003

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Anthony, R. G., M. A. O'Connell, M. M. Pollock, and J. G. Hallett. 2003. Associations of mammals with riparian ecosystems in Pacific Northwest forests. Pages 510–563 in C. J. Zabel and R. G. Anthony, editors. Mammal Community Dynamics. Management and Conservation in the Coniferous Forests of Western North America. Cambridge University Press, Cambridge, UK
Arbogast, B. S. 1999a. Mitochondrial DNA phylogeography of the New World flying squirrels (Glaucomys): implications for Pleistocene biogeography. Journal of Mammalogy 80:142–155CrossRefGoogle Scholar
Arbogast, B. S. 1999b. Comparative phylogeography of North American boreal mammals. PhD Dissertation, Wake Forest University, Winston-Salem, North Carolina, USA
Arbogast, B. S., and , G. J. Kenagy. 2001. Comparative phylogeography as an integrative approach to historical biogeography. Journal of Biogeography 28:819–825CrossRefGoogle Scholar
Arbogast, B. S., , R. A. Browne, and , P. D. Weigl. 2001. Evolutionary genetics and Pleistocene biogeography of North American tree squirrels (Tamiasciurus). Journal of Mammalogy 82:302–3192.0.CO;2>CrossRefGoogle Scholar
Armstrong, D. M. 1972. Distribution of mammals in Colorado. Volume 3, Monograph of the Museum of Natural History, University of Kansas, 415 pp
Aubry, K. B., M. J. Crites, and S. D. West. 1991. Regional patterns of small mammal abundance and community composition in Oregon and Washington. Pages 285–303 in L. F. Ruggiero et al., technical coordinators. Wildlife and vegetation of unmanaged Douglas-fir forests. USDA Forest Service General Technical Report PNW-285:1–533, Portland, Oregon, USA
Aubry, K. B., J. P. Hayes, B. L. Biswell, and B. G. Marcot. 2003. The ecological role of tree-dwelling mammals in western coniferous forests. Pages 405–443 in C. J. Zabel and R. G. Anthony, editors. Mammal Community Dynamics. Management and Conservation in the Coniferous Forests of Western North America. Cambridge University Press, Cambridge, UK
Banfield, A. W. F. 1974. The Mammals of Canada. University of Toronto Press, Toronto, Ontario, Canada
Barclay, R. M. R., and R. M. Brigham (editors). 1996. Bats and forests symposium, October 19–21, 1995, Victoria, British Columbia, Canada. Research Branch, British Columbia Ministry of Forests, Working Paper 23/1996:1–292
Brown, J. H. 1971a. Mechanisms of competitive exclusion between two species of chipmunks (Eutamias). Ecology 52:306–311CrossRefGoogle Scholar
Brown, J. H. 1971b. Mammals on mountaintops: non-equilibrium insular biogeography. American Naturalist 105:467–478CrossRefGoogle Scholar
Brown, J. H. 1978. The theory of insular biogeography and the distribution of boreal birds and mammals. The Great Basin Naturalist Memoirs 2:209–227Google Scholar
Buskirk, S. W. and W. J. Zielinksi. 2003. Small and mid-sized cornivores. Pages 207–249 in C. J. Zabel and R. G. Anthony, editors. Mammal Community Dynamics. Management and Conservation in the Coniferous Forests of Western North America. Cambridge University Press, Cambridge, UK
Byun, S. A., , B. F. Koop, and , T. E. Reichmen. 1997. North American black bear mtDNA phylogeography: implications for morphology and the Haida Gwaii glacial refugium controversy. Evolution 51:1647–1653Google ScholarPubMed
Carey, A. B. 1991. The biology of arboreal rodents in Douglas-fir forests. USDA Forest Service General Technical Report PNW-276:223–250, Portland, Oregon, USA
Carey, A. B. 1995. Sciurids in Pacific northwest managed and old-growth forest. Ecological Applications 5:648–661CrossRefGoogle Scholar
Carr, S. M., and S. A. Hicks. 1997. Are there two species of marten in North America? Genetic and evolutionary relationships within Martes. Pages 15–28 in G. Proulx, H. N. Bryant, and P. M. Woodard, editors. Martes: Taxonomy, Ecology, Techniques, and Management. Provincial Museum of Alberta, Canada
Chappell, M. A. 1978. Behavioral factors in the altitudinal zonation of chipmunks (Eutamias). Ecology 59:565–579CrossRefGoogle Scholar
Corn, P. S., and R. B. Bury. 1991. Small mammal communities in the Oregon Coast Range. Pages 241–254 in L. F. Ruggiero et al., technical coordinators. Wildlife and vegetation of unmanaged Douglas-fir forests. USDA Forest Service General Technical Report PNW-285:1–533, Portland, Oregon, USA
Corn, P. S., R. B. Bury, and T. A. Spies. 1988. Douglas-fir forests in the Cascade Mountains of Oregon and Washington: is the abundance of small mammals related to stand age and moisture? Pages 340–352 in R. C. Szaro et al., technical coordinators. Management of amphibians, reptiles, and small mammals in North America. USDA Forest Service General Technical Report RM-166:1–458, Flagstaff, Arizona, USA
Cowan, I. M. 1973. The mammals of British Columbia. Fifth edition. Volume 11, Handbook of the British Columbia Provincial Museum, Victoria, 414 pp
Cronin, M. A. 1992. Intraspecific variation in mitochondrial DNA of North American cervids. Journal of Mammalogy 73:70–82CrossRefGoogle Scholar
Dalquest, W. W. 1948. Mammals of Washington. Volume 2, Monograph of the Museum of Natural History, University of Kansas, 144 pp
Davis, R., and , C. Dunford. 1987. An example of contemporary colonization of montane islands by small, nonflying mammals in the American Southwest. American Naturalist 129:398–406CrossRefGoogle Scholar
Davis, R., , C. Dunford, and , M. V. Lomolino. 1988. Montane mammals of the American Southwest: the possible influence of post-Pleistocene colonization. Journal of Biogeography 15:841–848CrossRefGoogle Scholar
Davis, W. B. 1939. The Recent Mammals of Idaho. The Caxton Printers, Ltd., Caldwell, Idaho, USA
Demboski, J. R. and , J. A. Cook. 2001. Phylogeography of the dusky shrew, Sorex monticolus (Insectivora, Soricidae): insight into deep and shallow history in northwestern North America. Molecular Ecology 10:1227–1240CrossRefGoogle ScholarPubMed
Demboski, J. R., , K. D. Stone, and , J. A. Cook. 1999. Further perspectives on the Haida Gwaii glacial refugium. Evolution 53:2008–2012CrossRefGoogle ScholarPubMed
Durrant, S. D. 1952. Mammals of Utah. Taxonomy and distribution. Volume 6, Monographs of the Museum of Natural History, University of Kansas, 549 pp
Fenton, M. B. 1990. The foraging behaviour and ecology of animal-eating bats. Canadian Journal of Zoology 68:411–422CrossRefGoogle Scholar
Findley, J. S., A. H. Harris, D. E. Wilson, and C. Jones. 1975. Mammals of New Mexico. University of New Mexico Press, Albuquerque, New Mexico, USA
Fitzgerald, J. P., C. A. Meaney, and D. M. Armstrong. 1994. Mammals of Colorado. Denver Museum of Natural History and University Press of Colorado, Niwot, Colorado, USA
Foresman, K. R. 2001. The Wild Mammals of Montana. Volume 12, Special Publication, American Society of Mammalogists, 278 pp
Franklin, J. F., and J. A. Fites-Kaufmann. 1996. Assessment of late-successional forests of the Sierra Nevada. Pages 627–656 in Sierra Nevada Ecosystem Project. Status of the Sierra Nevada. Volume II, Assessments and scientific basis for management options. Volume 37, Wildland Resources Center Report, Centers for Water and Wildland Resources, University of California, Davis, 1528 pp
Gavin, T. A., , P. W. Sherman, , E. Yensen, and , B. May. 1999. Population genetic structure of the northern Idaho ground squirrel Spermophilus brunneus brunneus. Journal of Mammalogy 80:156–168CrossRefGoogle Scholar
Gilbert, F. F., and R. Allwine. 1991. Small mammal communities in the Oregon Cascade Range. Pages 257–267 in L. F. Ruggiero et al., technical coordinators. Wildlife and vegetation of unmanaged Douglas-fir forests. USDA Forest Service General Technical Report PNW-285:1–533, Portland, Oregon, USA
Graham, R. W. 1986. Response of mammalian communities to environmental changes during the Late Quaternary. Pages 300–313 in J. R. Diamond and T. J. Case, editors. Community Ecology. Harper and Row, New York, New York, USA
Graham, R. W., , E. L. Lundelius Jr., , M. A. Graham, , E. K. Schroeder, , R. S. Toomey III, , E. Anderson, , A. D. Barnosky, , J. A. Burns, , C. S. Churcher, , C. K. Grayson, , R. D. Guthrie, , C. R. Harrington, , G. T. Jefferson, , L. D. Martin, , H. G. McDonald, , R. E. Morlan, , H. A. Semken Jr., , S. D. Webb, , L. Werdelin, and , M. C. Wilson. 1996. Spatial response of mammals to late-Quaternary environmental fluctuations. Science 272:1601–1606Google ScholarPubMed
Grayson, D. 1993. The Deserts Past: A Natural Prehistory of the Great Basin. Smithsonian Institution Press, Washington, DC, USA
Grayson, D., and , D. B. Madsen. 2000. Biogeographic implications of recent low-elevation recolonization by Neotoma cinerea in the Great Basin. Journal of Mammalogy 81:1100–11052.0.CO;2>CrossRefGoogle Scholar
Grayson, D., , S. D. Livingston, , E. Rickart, and , M. W. Shaver III. 1996. Biogeographic significance of low elevation records for Neotoma cinerea from the northern Bonneville Basin, Utah. The Great Basin Naturalist 56:191–196Google Scholar
Hall, E. R. 1946. Mammals of Nevada. University of California Press, Berkeley, California, USA
Hall, E. R. 1981. The Mammals of North America. Two Volumes. John Wiley & Sons, New York, New York, USA
Hall, E. R. 1995. Mammals of Nevada. Second Edition. University of Nevada Press, Reno, Nevada, USA
Hallett, J. G., M. A. O'Connell, and C. C. Maguire. 2003. Ecological relationships of terrestrial small mammals in western coniferous forests. Pages 120–156 in C. J. Zabel and R. G. Anthony, editors. Mammal Community Dynamics. Management and Conservation in the Coniferous Forests of Western North America. Cambridge University Press, Cambridge, UK
Hamilton, W. J. III. 1962. Reproductive adaptations of the red tree mouse. Journal of Mammalogy 43:486–504CrossRefGoogle Scholar
Hayes, J. P. 2003. Habitat ecology and conservation of bats in western coniferous forests. Pages 81–119 in C. J. Zabel and R. G. Anthony, editors. Mammal Community Dynamics. Management and Conservation in the Coniferous Forests of Western North America. Cambridge University Press, Cambridge, UK
Hemstrom, M. A. 2003. Forests and woodlands of western North America. Pages 9–40 in C. J. Zabel and R. G. Anthony, editors. Mammal Community Dynamics. Management and Conservation in the Coniferous Forests of Western North America. Cambridge University Press, Cambridge, UK
Hoffmeister, D. F. 1986. Mammals of Arizona. University of Arizona Press, Tucson, Arizona, USA
Hogan, K. M., , M. C. Hedin, , H. S. Koh, , S. K. Davis, and , I. F. Greenbaum. 1993. Systematic and taxonomic implications of karyotypic, electrophoretic, and mitochondrial-DNA variation in Peromyscus from the Pacific Northwest. Journal of Mammalogy 74:819–830CrossRefGoogle Scholar
Holt, R. D., , G. R. Robinson, and , M. S. Gaines. 1995. Vegetation dynamics in an experimentally fragmented landscape. Ecology 76:1610–1624CrossRefGoogle Scholar
Ingles, L. G. 1948. Mammals of California. Stanford University Press, Stanford, California, USA
Ingles, L. G. 1954. Mammals of California and Its Coastal Waters. Stanford University Press, Stanford, California, USA
Ingles, L. G. 1965. Mammals of the Pacific States. California, Oregon, Washington. Stanford University Press, Stanford, California, USA
Kie, J. G., R. T. Bowyer, and K. M. Stewart. 2003. Ungulates in western coniferous forests: habitat relationships, population dynamics, and ecosystem processes. Pages 296–340 in C. J. Zabel and R. G. Anthony, editors. Mammal Community Dynamics. Management and Conservation in the Coniferous Forests of Western North America. Cambridge University Press, Cambridge, UK
Kirkland, G. L. Jr. 1985. Small mammal communities in temperate North American forests. Australian Mammalogy 8:137–144Google Scholar
Kunkel, K. E. 2003. Ecology, conservation, and restoration of large carnivores in western North America. Pages 250–295 in C. J. Zabel and R. G. Anthony, editors. Mammal Community Dynamics. Management and Conservation in the Coniferous Forests of Western North America. Cambridge University Press, Cambridge, UK
Kunz, T. H. 1982. Roosting ecology of bats. Pages 1–56 in T. H. Kunz, editor. Ecology of Bats. Plenum Press, New York, New York, USA
Kurtèn, B., and E. Anderson. 1980. Pleistocene Mammals of North America. Columbia University Press, New York, New York, USA
Larrison, E. J. 1976. Mammals of the Northwest. Washington, Oregon, Idaho, and British Columbia. Seattle Audubon Society, Seattle, Washington, USA
Larrison, E. J. and D. R. Johnson. 1981. Mammals of Idaho. University of Idaho Press, Pocatello, Idaho, USA
Lawlor, T. E. 1998. Biogeography of Great Basin mammals: paradigm lost?Journal of Mammalogy 79:1111–1130CrossRefGoogle Scholar
Lomolino, M. V., and , D. R. Perault. 2000. Assembly and disassembly of mammal communities in a fragmented temperate rain forest. Ecology 81:1517–1532Google Scholar
Lomolino, M. V., , J. H. Brown, and , R. Davis. 1989. Island biogeography of montane forest mammals in the American Southwest. Ecology 70:180–194CrossRefGoogle Scholar
Long, C. A. 1965. Mammals of Wyoming. Volume 14, Monograph of the Museum of Natural History, University of Kansas, pp. 493–758
MacMahon, J. A., , R. R. Parmenter, , K. A. Johnson, and , C. M. Crisafulli. 1989. Small mammal recolonization on the Mount St. Helens volcano: 1980–1987. American Midland Naturalist 122:365–387CrossRefGoogle Scholar
Martin, K. J., and W. C. McComb. 2003. Small mammals in a landscape mosaic: implications for conservation. Pages 567–586 in C. J. Zabel and R. G. Anthony, editors. Mammal Community Dynamics. Management and Conservation in the Coniferous Forests of Western North America. Cambridge University Press, Cambridge, UK
Maser, C. 1998. Mammals of the Pacific Northwest from the Coast to the High Cascades. Oregon State University Press, Corvallis, Oregon, USA
Maser, C., B. R. Mate, J. F. Franklin, and C. T. Dryness. 1981. Natural history of Oregon coast mammals. USDA Forest Service General Technical Report PNW-133:1–496, Portland, Oregon, USA
McNab, B. K. 1978. Energetics of arboreal folivores: physiological problems and ecological consequences of feeding on an ubiquitous food supply. Pages 153–162 in G. G. Montgomery, editor. The Ecology of Arboreal Folivores. Smithsonian Institution Press, Washington, DC, USA
Meiselman, N. 1992. Nest-site characteristics of red tree voles in Douglas-fir forests of northern California. Master's Thesis, Humboldt State University, Arcata, California, USA
Meiselman, N. and , A. T. Doyle. 1996. Habitat and microhabitat use by the red tree vole Phenacomys longicaudus. American Midland Naturalist 135:33–42CrossRefGoogle Scholar
Patterson, B. D. 1984. Mammalian extinction and biogeography in the southern Rocky Mountains. Pages 247–293 in M. H. Nitecki, editor. Extinctions. University of Chicago Press, Chicago, Illinois, USA
Patterson, B. D. 1995. Local extinctions and the biogeographic dynamics of boreal mammals in the Southwest. Pages 151–176 in C. Istock and R. S. Hoffman, editors. Storm Over a Mountain: Conservation Biology and the Mount Graham Affair. University of Arizona Press, Tucson, Arizona, USA
Perault, D. R. and , M. V. Lomolino. 2000. Corridors and mammal community structure across a fragmented, old-growth forest landscape. Ecological Monographs 70:401–422CrossRefGoogle Scholar
Ransome, D. B. and , T. P. Sullivan. 1997. Food limitation and habitat preference of Glaucomys sabrinus and Tamiasciurus hudsonicus. Journal of Mammalogy 78:538–549CrossRefGoogle Scholar
Raphael, M. G. 1988a. Long-term trends in abundance of amphibians, reptiles, and mammals in Douglas-fir forests of northwestern California. Pages 23–31 in R. C. Szaro et al., technical coordinators. Management of amphibians, reptiles, and small mammals in North America. USDA Forest Service General Technical Report RM-166:1–458, Flagstaff, Arizona, USA
Raphael, M. G. 1988b. Habitat associations of small mammals in a subalpine forest, Wyoming. Pages 359–367 in R. C. Szaro et al., technical coordinators. Management of amphibians, reptiles, and small mammals in North America. USDA Forest Service General Technical Report RM-166:1–458, Flagstaff, Arizona, USA
Raphael, M. G. 1991. Vertebrate species richness within and among seral stages of Douglas-fir/hardwood forest in northwestern California. Pages 415–423 in L. F. Ruggiero et al., technical coordinators. Wildlife and vegetation of unmanaged Douglas-fir forests. USDA Forest Service General Technical Report PNW-285:1–533, Portland, Oregon, USA
Rickart, E. A. 2001. Elevational diversity gradients, biogeography and structure of montane mammal communities in the intermountain region of North America. Global Ecology and Biogeography 10:77–100CrossRefGoogle Scholar
Ruggiero, L. F., K. B. Aubry, S. W. Buskirk, L. J. Lyon, and W. J. Zielinski, technical editors. 1994. The scientific basis for conserving forest carnivores. American marten, fisher, lynx, and wolverine in the western United States. USDA Forest Service General Technical Report RM-254:1–184, Fort Collins, Colorado, USA
Schweiger, E. W., , J. E. Diffendorfer, , R. D. Holt, , R. Pierotti, and , M. S. Gaines. 2000. The interaction of habitat fragmentation, plant, and small mammal succession in an old field. Ecological Monographs 70:383–400CrossRefGoogle Scholar
Smith, C. C. 1981. The indivisible niche of Tamiasciurus: an example of nonpartitioning of resources. Ecological Monographs 51:343–363CrossRefGoogle Scholar
Smith, H. C. 1993. Alberta Mammals. Atlas and Guide. Provincial Museum of Alberta, Edmonton, Alberta, Canada
Smith, W. P., R. G. Anthony, J. R. Waters, N. L. Dodd, and C. J. Zabel. 2003. Ecology and conservation of arboreal rodents of western coniferous forests. Pages 157–206 in C. J. Zabel and R. G. Anthony, editors. Mammal Community Dynamics. Management and Conservation in the Coniferous Forests of Western North America. Cambridge University Press, Cambridge, UK
Spies, T. A., and M. G. Turner. 1999. Dynamic forest mosaics. Pages 95–160 in M. L. Hunter, Jr., editor. Maintaining Biodiversity in Forest Ecosystems. Cambridge University Press, Cambridge, UK
Stone, K. D., and , J. A. Cook. 2000. Phylogeography of black bears (Ursus americanus) of the Pacific Northwest. Canadian Journal of Zoology 78:1218–1223CrossRefGoogle Scholar
Sutton, D. A., and , C. F. Nadler. 1974. Systematic revision of three Townsend chipmunks (Eutamias townsendii). Southwestern Naturalist 19:199–212CrossRefGoogle Scholar
Sutton, D. A., and , B. D. Patterson. 2000. Geographic variation of the western chipmunks Tamias senex and T. siskiyou, with two new subspecies from California. Journal of Mammalogy 81:299–3162.0.CO;2>CrossRefGoogle Scholar
Thompson, J. L., and , L. V. Diller. 2002. Relative abundance, nest site characteristics, and nest dynamics of dusky tree voles on managed timberlands in coastal northwest California. Northwestern Naturalist 83:91–100CrossRefGoogle Scholar
Vaughan, T. A., and , N. J. Czaplewski. 1985. Reproduction in Stephen's woodrat: the wages of folivory. Journal of Mammalogy 66:429–443CrossRefGoogle Scholar
Verts, B. J., and , L. N. Carraway. 1995. Phenacomys albipes. Mammalian Species 494:1–5CrossRefGoogle Scholar
Verts, B. J., and L. N. Carraway. 1998. Land Mammals of Oregon. University of California Press, Berkeley, California, USA
Wilson, D. E., and D. M. Reeder, editors. 1993. Mammal Species of the World. A Taxonomic and Geographic Reference. Smithsonian Institution Press, Washington, DC, USA
Wilson, D. E., and S. Ruff, editors. 1999. The Smithsonian Book of North American Mammals. Smithsonian Institution Press, Washington, DC, USA
Wooding, S., and , R. Ward. 1997. Phylogeography and Pleistocene evolution in the North American black bear. Molecular Biology and Evolution 14:1096–1105CrossRefGoogle ScholarPubMed
Zeveloff, S. I., and F. R. Collett. 1988. Mammals of the Intermountain West. University of Utah Press, Salt Lake City, Utah, USA

Save book to Kindle

To save this book to your Kindle, first ensure coreplatform@cambridge.org is added to your Approved Personal Document E-mail List under your Personal Document Settings on the Manage Your Content and Devices page of your Amazon account. Then enter the ‘name’ part of your Kindle email address below. Find out more about saving to your Kindle.

Note you can select to save to either the @free.kindle.com or @kindle.com variations. ‘@free.kindle.com’ emails are free but can only be saved to your device when it is connected to wi-fi. ‘@kindle.com’ emails can be delivered even when you are not connected to wi-fi, but note that service fees apply.

Find out more about the Kindle Personal Document Service.

Available formats
×

Save book to Dropbox

To save content items to your account, please confirm that you agree to abide by our usage policies. If this is the first time you use this feature, you will be asked to authorise Cambridge Core to connect with your account. Find out more about saving content to Dropbox.

Available formats
×

Save book to Google Drive

To save content items to your account, please confirm that you agree to abide by our usage policies. If this is the first time you use this feature, you will be asked to authorise Cambridge Core to connect with your account. Find out more about saving content to Google Drive.

Available formats
×