Hostname: page-component-78c5997874-fbnjt Total loading time: 0 Render date: 2024-11-17T19:32:52.712Z Has data issue: false hasContentIssue false

Middle Cambrian fossils from Tizi N'Tichka, the High Atlas, Morocco. Part 2. Hyolitha

Published online by Cambridge University Press:  14 July 2015

Ladislav Marek
Affiliation:
Geologicky ústav AVCR, Rozvojová 135, 16500 Prague 6, Czech Republic
John M. Malinky
Affiliation:
Academy of Science, Geological Institute, Rozvojova 135, 16502 Prague 6, Czech Republic
Gerd Geyer
Affiliation:
Institut für Paläontologie, Pleicherwall 1, D-97070, Würzburg, Federal Republic of Germany

Abstract

Shelly fossils from the basal Middle Cambrian, near the peak of Tizi n'Tichka in the central High Atlas, Morocco, include the new hyolithids Oboedalites peregrinus new species, Emarites decorissimus new genus and species, Patholites insolitus new genus and species, Rodentilites cricetus new genus and species, Slapylites afer new species, Tajinella mosei new genus and species, and Tizilites creber new genus and species. New orthothecid hyoliths are Gracilitheca destombesi new species, Contitheca chrosniaki new species, Sysoievia exilis new genus and species and an indeterminate species of Circotheca. The presence of Obeodalites, Slapylites, and Contitheca suggests a strong affinity to other Mediterranean and to Avalonian faunas. Their occurrence in Morocco extends their geographic ranges from Europe, and further reenforces their usefulness as indicators of Middle Cambrian age.

Type
Research Article
Copyright
Copyright © The Paleontological Society 

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Babcock, L. E., and Robison, R. A. 1988. Taxonomy and paleobiology of some Middle Cambrian Scenella (Cnidaria) and hyolithids (Mollusca) from western North America. University of Kansas Paleontological Contributions, Paper 121, 22 p.Google Scholar
Barrande, J. 1867. Système Silurien du Centre de la Bohême. Ordre des Pteropodes, 3. Prague, Paris, 179 p. (published by the author)Google Scholar
Bengtson, S., Conway Morris, S., Cooper, B. J., Jell, P. A., and Runnegar, B. N. 1990. Early Cambrian fossils from South Australia. Association of Australasian Paleontologists Memoir, 9, 364 p.Google Scholar
Billings, E. 1872. On some new fossils from the primordial rocks of Newfoundland. Canadian Naturalist, 6:465479.Google Scholar
Cobbold, E. S., and Pocock, R. W. 1934. The Cambrian area of Rushton (Shropshire). Philosophical Transactions of the Royal Society of London, Series B, 223:305409.Google Scholar
Dzik, J. 1981. Origin of the Cephalopoda. Acta Palaeontologica Polonica, 26(2):161191.Google Scholar
Eichwald, K. E. V. 1840. Über das silurische Schichtensystem in Esthland. Zeitschrift für Natur-und Heilkunde, St. Petersburg, Volumes 1 and 2, 210 p.CrossRefGoogle Scholar
Fisher, D. W. 1962. Small conoidal shells of uncertain affinities. In Moore, R. C. (ed.), Treatise on Invertebrate Paleontology. Part W, Miscellanea, W98-W143. Geological Society of America and University of Kansas Press, Lawrence, Kansas.Google Scholar
Grabau, A. W., and Shimer, H. W. 1910. North American index fossils; Invertebrates. Volume 2, A. G. Seilacher and Co., New York, 909 p.Google Scholar
Geyer, G. 1986. Mittelkambrische Mollusken aus Marokko und Spanien. Senckenbergiana lethaea, 67(1/4):55118.Google Scholar
Geyer, G. 1990a. Die marokkanischen Ellipsocephalidae (Trilobita: Redlichiida). Beringeria 3, 363 p.Google Scholar
Geyer, G. 1990b. Correlation along the Lower/Middle Cambrian boundary—a puzzling story with an elusory end? In Repina, L. N. and Zhuravlev, A. Yu., (eds.), Tretij mezhdunarodnyj simpozium po kembrijskoj sisteme. 1-9 avgusta 1990 g., SSSR, g. Novosibirsk. Tezisy Dokladov:100102.Google Scholar
Geyer, G., and Malinky, J. M. 1997. Middle Cambrian fossils from Tizi n'Tichka, the High Atlas, Morocco. Part 1. Introduction and trilobites. Journal of Paleontology, 71:620637.CrossRefGoogle Scholar
Geyer, G., Landing, E., and Malinky, J. 19XX. Middle Cambrian fossils from Tizi n'Tichka, the High Atlas, Morocco. Part 3. Small shelly fossils. Journal of Paleontology.Google Scholar
Hagdorn, H. 1995. Farbmuster und Pseudoskulptur bei Muschelkalkfossilien. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 195:85108.CrossRefGoogle Scholar
Hinz, I. 1987. The Lower Cambrian microfauna of Comley and Rushton, Shropshire, England. Palaeontographica, Abteilung A, 198:41100.Google Scholar
Holm, G. 1893. Sveriges kambrisk-siluriska Hyolithidae och Conulariidae. Sveriges geologiska Undersökning, Afhandlingar och Uppsatser, C 112, 172 p.Google Scholar
Houbrick, R. S., Stürmer, W., and Yochelson, E. L. 1988. Rare Mollusca from the Lower Devonian Hunsrück Slate of southern Germany. Lethaia, 21:395402.CrossRefGoogle Scholar
Hupé, P. 1959. Nouvelle contribution à l'étude du Cambrien marocain. Unpublished thesis 3éme cycle, Faculté des Sciences, Université de Paris, 447 p.Google Scholar
Landing, E. 1988. Lower Cambrian of eastern Massachusetts: Stratigraphy and small shelly fossils. Journal of Paleontology, 62:661695.Google Scholar
Malinky, J. M. 1988. Early Paleozoic Hyolitha from North America; reexamination of Walcott's and Resser's type specimens. Journal of Paleontology, 62:218233.CrossRefGoogle Scholar
Malinky, J. M. 1989. New Early Paleozoic Hyolithida and Orthothecida (Hyolitha) from North America. Journal of Paleontology, 63:302319.CrossRefGoogle Scholar
Malinky, J. M. 1990a. Early and Middle Cambrian Hyolitha (Mollusca) from northeastern China. Journal of Paleontology, 64:228239.CrossRefGoogle Scholar
Malinky, J. M. 1990b. Solenotheca, a new Hyolitha (Mollusca) from the Ordovician dovician of North America. Proceedings of the Biological Society of Washington, 103:265278.Google Scholar
Malinky, J. M., and Mapes, R. H. 1983. First occurrences of Hyolitha (Mollusca) in the Pennsylvanian of North America. Journal of Paleontology, 57:347353.Google Scholar
Malinky, J. M., Linsley, R. M., and Yochelson, E. L. 1987. Taxonomic revision of Hyolitha from the Middle Paleozoic of North America. Journal of Paleontology, 61:11731186.CrossRefGoogle Scholar
Malinky, J. M., and Mapes, R. H. 1983. First occurrences of Hyolitha (Mollusca) in the Pennsylvanian of North America. Journal of Paleontology, 57:347353.Google Scholar
Malinky, J. M., Mapes, R. H., and Broadhead, T. W. 1986. New Late Palaeozoic Hyolitha (Mollusca) from Oklahoma and Texas, and their palaeoenvironmental significance. Palaeontology, 29:303312.Google Scholar
Marek, L. 1963. New knowledge on the morphology of Hyolithes. Sborník geologickych ved, Rada Paleontologie, 1:5373.Google Scholar
Marek, L. 1966. New hyolithid genera from the Ordovician of Bohemia. Casopis Národního Muzea, 135:8992.Google Scholar
Marek, L. 1967. The class Hyolitha in the Caradoc of Bohemia. Sborník geologickych ved, Rada Paleontologie, 9:51113.Google Scholar
Marek, L. 1974. New Lower Ordovician hyolithid genera. Vestník Ustredního Ústavu Geologického, 49:287292.Google Scholar
Marek, L. 1976. On the ontogeny in Hyolithida. Casopis pro Mineralogii a Geologii, 21:277284.Google Scholar
Marek, L. 1980. Slapylites gen. n. from the Middle Cambrian of Bohemia (Hyolitha). Casopis Národního Muzea, 149:156160.Google Scholar
Marek, L. 1981. Middle Cambrian hyolithid family Parentilitidae fam. nov. Casopis Národního Muzea, 150:163168.Google Scholar
Marek, L. 1983. The Ordovician hyoliths of Anti-Atlas (Morocco). Sborník Národního Muzea v Praze, 39:136.Google Scholar
Marek, L., and Yochelson, E. L. 1962. Paleozoic mollusk: Hyolithes. Science, 146:16741675.CrossRefGoogle Scholar
Marek, L., and Yochelson, E. L. 1976. Aspects of the biology of Hyolitha (Mollusca). Lethaia 9:6582.CrossRefGoogle Scholar
Matthew, G. F. 1900. Preliminary notice of the Etcheminian fauna of Cape Breton. Bulletin of Natural History Society of New Brunswick, 18:198208.Google Scholar
Meshkova, N. P. 1974. Khiolity nizhnego kembriya Sibirskoj platformy [Hyoliths of the Lower Cambrian of the Siberian Platform]. Trudy Institut geologii geofiziki, Sibirskoe Otdelenie. Nauka Publishers, Novosibirsk, 97, 110 p.Google Scholar
Missarzhevskij, V. V. 1969. Opisanie khiolitov, gastropod, khiolitel'mintov, kamenid i form neyasnogo sistematicheskogo polozheniya [Descriptions of hyoliths, gastropods, hyolithelminths, camenids, and forms of an obscure systematic position]. In Rozanov, A. Yu., Missarzhevskij, V. V.Volkova, N. A., Voronova, L. G., Krylov, I. N., Keller, B. M., Korolyuk, I. K., Lendzion, K., Mikhnyak, R., Pykhova, N. G. & Sidorov, A. D., Tommotskij yarus i problema nizhnej granitsy Kembriya. Akademiya Nauk SSSR, Ordena Trudovogo krasnogo znameni Geologicheskij Institut, Trudy, 206:105175, Moscow (Nauka).Google Scholar
Missarzhevskij, V. V. 1989. Drevnejskie skeletnye okamenelosti i stratigrafiya pogranichnykh tolshch dokembriya i kembriya [Oldest skeletal fossils and stratigraphy of Precambrian and Cambrian boundary beds]. Trudy Ordena Trudov, Krasnogo Zhameni geologicheskij Institut Akademiya Nauk SSSR, 443:238 p., Nauka (Moscow).Google Scholar
Neltner, L. 1938. Etudes géologique dans le Sud marocain (Haut Atlas et Anti-Atlas). Notes et Mémoirs de la Service des Mines et Carte géologique de Maroc, 42, 298 p.Google Scholar
Novak, O. 1891. Revision der paläozoischen Hyolithiden Böhmens. Abhandlungen der mathematisch-naturwisschenschaftlichen Classe der Böhmischen Gesellschaft der Wissenschaften, Series 7, 4, 48 p.Google Scholar
Pojeta, J. Jr. 1987. Phylum Hyolitha, p. 436444. In Boardman, R. S., Cheetham, A. H., and Rowell, A. J. (eds.), Fossil Invertebrates. Blackwell Scientific Publications, Palo Alto.Google Scholar
Pojeta, J. Jr., and Gordon, M. Jr. 1987. Class Cephalopoda, p. 329358. In Boardman, R. S., Cheetham, A. H., and Rowell, A. J. (eds.), Fossil Invertebrates. Blackwell Scientific Publications, Palo Alto.Google Scholar
Poulsen, C. 1932. The Lower Cambrian faunas of East Greenland. Meddelelser om Gronland, 87(6), 66 p.Google Scholar
Yi, Qian, 1989. Early Cambrian small shelly fossils of China with special reference to the Precambrian-Cambrian boundary. Nanjing University Publishing House, Nanjing, 341 p. (In English)Google Scholar
Yi, Qian, and Bengtson, S. 1989. Palaeontology and biostratigraphy of the Early Cambrian Meishucunian Stage in Yunnan Province, South China. Fossils and Strata 24, 156 p.Google Scholar
Runnegar, B. 1980. Hyolitha: status of the phylum. Lethaia, 13:2125.CrossRefGoogle Scholar
Runnegar, B., Pojeta, J. Jr, Morris, N. I., Taylor, J. D., Taylor, M. E., and McClung, G. 1975. Biology of Hyolitha. Lethaia, 7:181191.CrossRefGoogle Scholar
Saito, K. 1936. Older Cambrian Brachiopoda, Gastropoda, etc. from North-western Korea. Journal of the Faculty of Science, University of Tokyo, Sect. 2, 4:345367.Google Scholar
Salter, J. W. 1866. On the fossils of North Wales. Memoirs of the Geological Survey of Great Britain, III, Appendix, 239381.Google Scholar
Shergold, J. H., Cooper, R. A., Mackinnon, D. I., and Yochelson, E. L. 1976. Late Cambrian Brachiopoda, Mollusca, and Trilobita from Northern Victoria Land, Antarctica. Palaeontology, 19:247291.Google Scholar
Sysoiev, V. A. 1958. Nadotryad Hyolithoida. In Osnovy paleontologii. Molluski-golovongie II. (Superorder Hyolithoides. Fundamentals of Paleontology. Mollusca, Part 2), Akademiya Nauk SSSR, 2:184190. (In Russian)Google Scholar
Sysoiev, V. A. 1959. Khiolity rodov Circotheca iz nizhnego kembriya Tajmirskogo natsional'nogo okruga (Hyolithids of the genus Circotheca from the Lower Cambrian of the Tajmyr National District). Paleontologicheskij Zhurnal, 1959:8494. (In Russian)Google Scholar
Sysoiev, V. A. 1962. Khiolity kembriya severnogo sklona Aldanskogo shchita (Cambrian hyolithids from the northern slope of the Aldan shield). Akademiya Nauk SSSR, Yakutskij filial Sibirskogo otdeleniya, Yakutsk, 65 p. (In Russian)Google Scholar
Sysoiev, V. A. 1968. Stratigrafiya i khiolity drevnejshikh sloev nizhnego kembriya Sibirskoj platformy (Stratigraphy and hyoliths of the oldest Lower Cambrian beds of the Siberian platform). Akademiya Nauk SSSR, Yakutskij filial Sibirskogo otdeleniya Instituta Geologii, Yakutsk, 67 p. (In Russian)Google Scholar
Sysoiev, V. A. 1972. Biostratigrafiya i khiolity ortotetsiomorfy nizhnego kembriya Sibirskoj platformy (Biostratigraphy and orthothecomorph hyoliths from the Lower Cambrian of the Siberian platform). Nauka Publishers, Moscow, 152 p. (In Russian)Google Scholar
Sysoiev, V. A. 1984. Morfologiya i sistematicheskoye polozheniye khiolitov. Paleontologicheskij Zhurnal, 1984(2):314(Morphology and systematic position of the hyoliths. Paleontological Journal, 1984:1-10).Google Scholar
Termier, G., and Termier, H., 1950. Paléontologie Marocaine II. Invertébrés de L'ère Primaire Fascicule IV. Annélides, Arthropodes, Echinodermes, Conularides et Graptolithes. Notes et Mémoirs de la Service des Mines et Carte géologique du Maroc, 79 p.Google Scholar
Walcott, C. D. 1884. Paleontology of the Eureka District, Nevada. U. S. Geological Survey, Monographs, 8, 298 p.Google Scholar
Walcott, C. D. 1890. Descriptive notes on new genera and species from the Lower Cambrian or Olenellus zone of North America. U. S. National Museum, Proceedings, 13 (for 1889):3346.CrossRefGoogle Scholar
Webers, G. F., Pojeta, J. Jr., and Yochelson, E. L., 1992. Cambrian mollusca from the Minaret Formation, Ellsworth Mountains, West Antarctica, p. 181248. In Webers, G. F., Craddock, C., and Splettstoesser, J. F. (eds.), Geology and Paleontology of the Ellsworth Mountains, West Antarctica. Geological Society of America Memoir 170, Boulder, Colorado.CrossRefGoogle Scholar
Yochelson, E. L. 1974. Redescription of the Early Cambrian Helenia bella Walcott, an appendage of Hyolithes. U. S. Geological Survey Journal of Research, 2(6):717722.Google Scholar
Yochelson, E. L. 1979. The radiation of Mollusca and mollusc-like groups, p. 323358. In House, M. R. (ed.), The Origin of Major Invertebrate Groups. Systematic Association Special Volume 12.Google Scholar