Fine roots and their turnover represent a dynamic aspect of below-ground biomass (BGB) and nutrient capital in forest ecosystems, and account for a significant fraction of net primary productivity (NPP) (Cuevas 1995, Vogt et al. 1990). On a weight basis, coarse roots contribute more to total ecosystem biomass than fine roots, but they account for only a small portion of annual root production (Eamus et al. 2002). Despite the fact that fine roots may compose less than 2% of total ecosystem biomass, they may contribute up to 40% of total ecosystem production (Vogt et al. 1990). Therefore, estimates of root production, like estimates of root biomass, should differentiate between coarse- and fine-root production.