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Was FLK North levels 1–2 a classic “living floor” of Oldowan hominins or a taphonomically complex palimpsest dominated by large carnivore feeding behavior?

Published online by Cambridge University Press:  20 January 2017

H.T. Bunn*
Affiliation:
Department of Anthropology, University of Wisconsin-Madison, 1180 Observatory Drive, Madison, WI 53706, USA
A.Z.P. Mabulla
Affiliation:
Archaeology Unit, University of Dar es Salaam, Dar es Salaam, P.O. Box 35050, Tanzania
M. Domínguez-Rodrigo
Affiliation:
IDEA (Instituto de Evolución en África), Museo de los Orígenes, Plaza de San Andrés 2, 28005 Madrid, Spain Department of Prehistory, Complutense University, Prof. Aranguren s/n, 28040, Madrid, Spain
G.M. Ashley
Affiliation:
Department of Geological Sciences, Rutgers University, 610 Taylor Road, Piscataway, NJ 08854, USA
R. Barba
Affiliation:
Department of Prehistory, Complutense University, Prof. Aranguren s/n, 28040, Madrid, Spain
F. Diez-Martín
Affiliation:
Department of Prehistory and Archaeology, University of Valladolid, Plaza del Campus s/n, 47011, Valladolid, Spain
K. Remer
Affiliation:
Department of Anthropology, University of Wisconsin-Madison, 1180 Observatory Drive, Madison, WI 53706, USA
J. Yravedra
Affiliation:
Department of Prehistory, Complutense University, Prof. Aranguren s/n, 28040, Madrid, Spain
E. Baquedano
Affiliation:
Museo Arqueológico Regional, Plaza de las Bernardas s/n, 28801 Alcalá de Henares, Madrid, Spain
*
*Corresponding author. E-mail address:htbunn@wisc.edu (H.T. Bunn).

Abstract

From excavation at FLK North levels 1–2 in 1960–1962, Mary Leakey reported approximately 1200 Oldowan artifacts and 3300 large mammal fossils as a hominin “living floor”. Preliminary taphonomic analysis by Bunn seemed supportive, based on the presence of some cut-marked bones, the concentration of several dozen bovid individuals, and the relative abundance of limbs and mandibles over other axial elements. Recent taphonomic analysis of Leakey's entire fossil assemblage by Domínguez-Rodrigo and Barba, however, documents a minor hominin role at the site, contrasted to the dominant role of carnivores. Felids brought prey animals; hyenas scavenged from abandoned felid meals. At different times, hominins butchered several bovids and discarded artifacts at this dynamic location. Since 2006, renewed excavations at FLK North and other sites by the Olduvai Paleoanthropology and Paleoecology Project (TOPPP) have expanded artifact and fossil samples and implemented new analytical approaches to clarify taphonomic histories of the Olduvai paleolandscape. At FLK North, > 1000 new large mammal fossils from levels 1 to 2 show minimal butchery evidence amid abundant evidence of carnivore gnawing/fracture, rodent gnawing, and sediment abrasion. To help guide future excavation and analyses, we have developed several alternative working hypotheses of site formation.

Type
Special Issue Articles
Copyright
University of Washington

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References

Ashley, G.M., Barboni, D., Domínguez-Rodrigo, M., Bunn, H.T., Mabulla, A.Z.P., Diez- Martín, F., Barba, R., and Baquedano, E. Paleoenvironmental and paleoecological reconstruction of a freshwater oasis in savannah grassland at FLK North, Olduvai Gorge, Tanzania. Quaternary Research 74, (2010). 333343.Google Scholar
Binford, L.R. Bones: Ancient Men and Modern Myths. (1981). Academic Press, New York.Google Scholar
Binford, L.R. Human ancestors: changing views of their behavior. Journal of Anthropological Archaeology 4, (1985). 292327.Google Scholar
Binford, L.R. Comment on Systematic butchery by Plio-Pleistocene hominids at Olduvai Gorge, Tanzania by H.T. Bunn and E.M. Kroll. Current Anthropology 27, (1986). 444446.Google Scholar
Binford, L.R. Fact and fiction about the Zinjanthropus floor: data, arguments, and interpretations. Current Anthropology 29, (1988). 123135.Google Scholar
Blumenschine, R.J. Early Hominid Scavenging Opportunities: Implications of Carcass Availability in the Serengeti and Ngorongoro Ecosystems. (1986). British Archaeological Reports, International Series. 283 Google Scholar
Blumenschine, R.J. Comment on Systematic butchery by Plio-Pleistocene hominids at Olduvai Gorge, Tanzania by H.T. Bunn and E.M. Kroll. Current Anthropology 27, (1986). 446 Google Scholar
Blumenschine, R.J. Characteristics of an early hominid scavenging niche. Current Anthropology 28, (1987). 383407.Google Scholar
Blumenschine, R.J., and Cavallo, J.A. Scavenging and human evolution. Scientific American 267, (1992). 9096.Google Scholar
Blumenschine, R.J., and Pobiner, B.L. Zooarchaeology and the ecology of Oldowan hominin carnivory. Ungar, P.S. Evolution of the Human Diet. (2007). Oxford University Press, New York. 167190.Google Scholar
Blumenschine, R.J., Peters, C.R., Capaldo, S.D., Andrews, P., Njau, J.K., and Pobiner, B.L. Vertebrate taphonomic perspectives on Oldowan hominin land use in the Plio- Pleistocene Olduvai Basin, Tanzania. Pickering, T.R., Schick, K., and Toth, N. Breathing Life into Fossils: Taphonomic Studies in Honor of C.K. (Bob) Brain. (2007). Stone Age Institute Press, Gosport, Indiana. 161179.Google Scholar
Blumenschine, R.J., Prassack, K.A., Kreger, C.D., and Pante, M.C. Carnivore tooth-marks, microbial bioerosion, and the invalidation of Domínguez-Rodrigo and Barba's (2006) test of Oldowan hominin scavenging behavior. Journal of Human Evolution 53, (2007). 420426.Google Scholar
Bunn, H.T. Archaeological evidence for meat-eating by Plio-Pleistocene hominids from Koobi Fora and Olduvai Gorge. Nature 291, (1981). 574577.Google Scholar
Bunn, H.T. Meat-eating and Human Evolution: Studies on the Diet and Subsistence Patterns of Plio-Pleistocene Hominids in East Africa. Ph.D. dissertation. (1982). University of California, Berkeley.Google Scholar
Bunn, H.T. Patterns of skeletal representation and hominid subsistence activities at Olduvai Gorge, Tanzania, and Koobi Fora, Kenya. Journal of Human Evolution 15, (1986). 673690.Google Scholar
Bunn, H.T. Bone assemblages at base camps: a further consideration of carcass transport and bone destruction by the Hadza. Hudson, J. From Bones to Behavior: Ethnoarchaeological and Experimental Contributions to the Interpretation of Faunal Remains. Occasional Paper No. 21. (1993). Center for Archaeological Investigations, Southern Illinois University, Carbondale, Illinois. 156168.Google Scholar
Bunn, H.T. Early Pleistocene hominid foraging strategies along the ancestral Omo river at Koobi Fora, Kenya. Journal of Human Evolution 27, (1994). 247266.Google Scholar
Bunn, H.T. The bone assemblages from the excavated sites. Isaac, G.Ll. Koobi Fora Project Monograph, Volume 5. (1997). Clarendon Press, Oxford. 402458.Google Scholar
Bunn, H.T. Butchering backstraps and bearing backbones: insights from Hadza foragers and implications for Paleolithic archaeology. Pickering, T.R., Schick, K., and Toth, N. Breathing Life into Fossils: Taphonomic Studies in Honor of C.K. (Bob) Brain. (2007). Stone Age Institute Press, Gosport, Indiana. 269279.Google Scholar
Bunn, H.T. Meat made us human. Ungar, P.S. Evolution of the Human Diet. (2007). Oxford University Press, New York. 191211.Google Scholar
Bunn, H.T., and Ezzo, J.A. Hunting and scavenging by Plio-Pleistocene hominids: nutritional constraints, archaeological patterns, and behavioral implications. Journal of Archaeological Science 20, (1993). 365398.Google Scholar
Bunn, H.T., and Kroll, E.M. Systematic butchery by Plio-Pleistocene hominids at Olduvai Gorge, Tanzania. Current Anthropology 27, (1986). 431452.CrossRefGoogle Scholar
Bunn, H.T., and Kroll, E.M. Fact and fiction about the FLK Zinjanthropus floor: data, arguments, and interpretations. (reply to L.R. Binford). Current Anthropology 29, (1988). 135149.Google Scholar
Bunn, H., Harris, J.W.K., Isaac, G., Kaufulu, Z., Kroll, E., Schick, K., Toth, N., and Behrensmeyer, A.K. FxJj 50: an early Pleistocene site in northern Kenya. World Archaeology 12, (1980). 109136.Google Scholar
Bunn, H.T., Bartram, L.E., and Kroll, E.M. Variability in bone assemblage formation from Hadza hunting, scavenging, and carcass processing. Journal of Anthropological Archaeology 7, (1988). 412457.Google Scholar
Cushing, A.E. The landscape zooarchaeology and paleontology of Plio-Pleistocene Olduvai, Tanzania, and their implications for early hominid ecology. Ph.D. dissertation. (2002). Rutgers University, New Brunswick.Google Scholar
Diez-Martín, F., Sanchez Yustos, P., Domínguez-Rodrigo, M., Mabulla, A.Z.P., Bunn, H.T., Ashley, G.M., Barba, R., and Baquedano, E. New insights into hominin lithic activities at FLK North Bed I, Olduvai Gorge, Tanzania. Quaternary Research 74, (2010). 376387.CrossRefGoogle Scholar
Domínguez-Rodrigo, M. Meat-eating by early hominids at the FLK 22 Zinjanthropus site, Olduvai Gorge, Tanzania: an experimental approach using cut mark data. Journal of Human Evolution 33, (1997). 669690.Google Scholar
Domínguez-Rodrigo, M. A reassessment of the study of cut mark patterns to infer hominid manipulation of fleshed carcasses at the FLK Zinj 22 site, Olduvai Gorge, Tanzania. Trabajos de Prehistoria 54, (1997). 2942.Google Scholar
Domínguez-Rodrigo, M. Meat-eating and carcass procurement by hominids at the FLK Zinj 22 site, Olduvai Gorge, Tanzania: a new experimental approach to the old hunting-versus-scavenging debate. Ullrich, H. Lifestyles and Survival Strategies in Pliocene and Pleistocene Hominids. (1999). Edition Archaea, Schwelm, Germany. 89111.Google Scholar
Domínguez-Rodrigo, M., and Barba, R. New estimates of tooth marks and percussion marks from FLK Zinj, Olduvai Gorge (Tanzania): the carnivore–hominid–carnivore hypothesis falsified. Journal of Human Evolution 50, (2006). 170194.Google Scholar
Domínguez-Rodrigo, M., and Barba, R. Five more arguments to invalidate the passive scavenging version of the carnivore–hominid–carnivore model: a reply to Blumenschine et al. (2007a). Journal of Human Evolution 53, (2007). 427433.Google Scholar
Domínguez-Rodrigo, M., and Barba, R. A palimpsest at FLK North 1-2: independent carnivore- and hominid-made bone accumulations. Domínguez-Rodrigo, M., Barba, R., and Egeland, C.P. Deconstructing Olduvai: A Taphonomic Study of the Bed I Sites. (2007). Springer, New York. 127163.Google Scholar
Domínguez-Rodrigo, M., and Barba, R. The behavioral meaning of cut marks at the FLK Zinj level: the carnivore–hominid–carnivore hypothesis falsified (II). Domínguez-Rodrigo, M., Barba, R., and Egeland, C.P. Deconstructing Olduvai: A Taphonomic Study of the Bed I Sites. (2007). Springer, New York. 75100.Google Scholar
Domínguez-Rodrigo, M., Barba, R., and Egeland, C.P. Deconstructing Olduvai: A Taphonomic Study of the Bed I Sites. (2007). Springer, New York.Google Scholar
Domínguez-Rodrigo, M., Bunn, H.T., Mabulla, A.Z., Ashley, G.M., Diez-Martín, F., Barba, R., and Baquedano, E. Disentangling hominin and carnivore activities near a spring at FLK North (Olduvai Gorge, Tanzania). Quaternary Research 74, (2010). 363375.Google Scholar
Isaac, G.Ll. The diet of early man: aspects of archaeological evidence from lower and middle Pleistocene sites in Africa. World Archaeology 2, (1971). 278299.Google Scholar
Isaac, G.Ll. The food-sharing behavior of protohuman hominids. Scientific American 238, (1978). 90108.Google Scholar
Isaac, G.Ll. Bones in contention: competing explanations for the juxtaposition of early Pleistocene artifacts and faunal remains. Clutton-Brock, J., and Grigson, C. Animals and Archaeology: Hunters and their Prey. British Archaeological Reports. International Series 163, (1983). 319.Google Scholar
Kroll, E.M., and Isaac, G.Ll. Configurations of artifacts and bones at early Pleistocene sites in East Africa. Hietela, H. Intrasite Spatial Analysis in Archaeology. (1984). Cambridge University Press, Cambridge. 431.Google Scholar
Leakey, M. Olduvai Gorge, vol. 3. Excavations in Bed I and II. (1971). Cambridge University Press, Cambridge. 1960–1963 Google Scholar
Pobiner, B.L., Rogers, M.J., Monahan, C.M., and Harris, J.W.K. New evidence for hominin carcass processing strategies at 1.5 Ma, Koobi Fora, Kenya. Journal of Human Evolution 55, (2008). 103130.Google Scholar
Potts, R. Home bases and early hominids. American Scientist 72, (1984). 338347.Google Scholar
Potts, R., and Shipman, P. Cutmarks made by stone tools on bones from Olduvai Gorge, Tanzania. Nature 291, (1981). 577580.Google Scholar