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New species of Liostracina Monke, 1903 (Trilobita, Cambrian) from Yunnan, China: complete holaspid exoskeleton and implications for higher level classification

Published online by Cambridge University Press:  11 July 2022

Xianfeng Yang ,
Shanchi Peng Open the ORCID record for Shanchi Peng [Opens in a new window] ,
Loren E. Babcock ,
Xuejian Zhu  and
Yu Liu
Xianfeng Yang
Affiliation:
Yunnan Key Laboratory for Palaeobiology, Institute of Palaeontology, Yunnan University, Kunming 650500, China MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Yunnan University, Kunming 650500, China
Shanchi Peng*
Affiliation:
State Key Laboratory of Palaeobiology and Stratigraphy, Chinese Academy of Sciences, Nanjing 210008, China Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China
Loren E. Babcock
Affiliation:
Orton Geological Museum, School of Earth Sciences, The Ohio State University, 125 South Oval Mall, Columbus, OH 43210, USA
Xuejian Zhu
Affiliation:
State Key Laboratory of Palaeobiology and Stratigraphy, Chinese Academy of Sciences, Nanjing 210008, China Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China
Yu Liu
Affiliation:
Yunnan Key Laboratory for Palaeobiology, Institute of Palaeontology, Yunnan University, Kunming 650500, China MEC International Joint Laboratory for Palaeobiology and Palaeoenvironment, Yunnan University, Kunming 650500, China
*
*Corresponding author.
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Abstract

The trilobite Liostracina has been recognized as important in taxonomic and stratigraphic studies for more than a century. Until now, the genus Liostracina and family Liostracinidae have been known from only incomplete holaspid material, a degree 2 meraspis, and protaspides. A new locality in the Longha Formation (Cambrian: Guzhangian) of southeastern Yunnan, China, yields a rich collection of articulated holaspid exoskeletons and disarticulated sclerites of a new species, Liostracina fuluensis n. sp. These specimens reveal thoracic and ventral morphology that was previously unknown for Liostracina; they demonstrate that it has a rostral plate plus rostral and connective sutures, rather than a ventral median suture on the cephalic doublure. This confirms a natant hypostomal condition for the genus. Holaspid exoskeletal features, combined with evidence of a non-asaphoid-type protaspis, indicate that Liostracina is neither a primitive trinucleoid nor a primitive asaphoid. The family Liostracinidae is excluded from the superfamily Trinucleoidea and the order Trinucleida and reassigned to the order Ptychopariida.

UUID: http://zoobank.org/48dcbe31-e25d-4657-8441-732d5ea2b49d

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Journal of Paleontology , Volume 96 , Issue 6 , November 2022 , pp. 1431 - 1446
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Copyright © The Author(s), 2022. Published by Cambridge University Press on behalf of The Paleontological Society

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