Drillholes on shells provide a useful way to investigate prey and predator relationships. The current study documents predator–prey interactions exemplified by a faunal assemblage of the fossil gastropod Turritella cingulifera from the Pleistocene Szekou Formation in Hengchun Peninsula, Taiwan. All recognisable skeletal and shell fragments that are larger than 3 mm in size were collected and recorded. Processed bulk sediments (5.24 kg) contained 1462 molluscan shells, including 824 specimens of T. cingulifera, and 27 non-molluscan invertebrates. In the current study, approximately 41.6% (609/1462) of molluscs are drilled with at least one hole. Drilling intensities (DIs) regardless of shell completeness in all gastropods, bivalves and the turritelline gastropod T. cingulifera are 0.546, 0.060 and 0.413, respectively. DI on turritellids is significantly lower than that on other gastropods (χ2= 21.039, P < 0.001). Furthermore, the percentage of drillholes that occur in multiply drilled specimens is 34.7% (95/275) for turritelline gastropods based on complete to nearly complete specimens (n = 588). Our study shows no significant preference of drillhole position either on the suture or on the whorl (χ2= 0.055, P = 0.814). Most drillholes are located in whorls two to four proximal to the aperture. Drillhole diameters of the shells with one drillhole and ones with multiple drillholes are 1.0 and 0.5 mm on average, and the results of Mann–Whitney tests indicate that they are significantly different (P < 0.001). The first turritelline gastropod shell with an incomplete drillhole from Taiwan is documented here. The dominant drilling predators were naticids based on the drillhole morphology and the presence of naticids in the same assemblage. No apparent prey size selectivity is observed, so a ‘size refugium’ does not exist for the turritellids in the current study.

Methodologies that analyse the colouration and external appearance of extant species are very useful tools when facing one of the greatest challenges in the palaeoartistic reconstructions of extinct fauna: inferring the colour patterns. Earlier works have applied this methodology, for example, in the reconstruction of the Miocene bovid Tethytragus, proving that the maximum likelihood (ML) analysis to infer ancestral states has promising potential. This study offers a proposal for the reconstruction of the external appearance of Heteroprox moralesi Azanza 1989, an early cervid of the Middle Miocene present in several fossil sites of Central Spain. For the reconstruction of the external appearance, the colour patterns of all the extant species of the family Cervidae were studied with the method of ML analysis, as well as recent works about their phylogeny. The results show the most probable basal colour pattern of the cervids: dark shades on the limbs, dorsal section, and head of the animal, and, in contrast, lighter colours on the neck and perianal region. This basal pattern can be used as a basis for reconstructing colouration and to hypothesise about the external appearance of extinct taxa. Furthermore, the inferred forest habitat of H. moralesi has been taken into consideration in order to adjust the colour pattern, comparing the final results of the analysis performed in this study with that of the pattern observed in extant forest deer as well as with previous works employing this methodology.

In the last decade, the identification of bone fragments by peptide mass fingerprinting or zooarchaeology by mass spectrometry is developing as a powerful tool in Quaternary palaeontology. The sequence of amino acids that make up the bone collagen molecule shows slight variations between taxa, which can be studied by mass spectrometry for taxonomic purposes. This requires reference databases that allow peptide identification. Although the cave bear (Ursus spelaeus Rosenmüller, 1794) is a common component in many European Pleistocene cave sites, no peptide fingerprint taxonomic study has paid special attention to this species up to now. For peptide markers in Ursidae, the most recent proposal is based on collagen obtained from a modern brown bear sample. In this work we attempt to cover this gap by studying bone collagen of cave and brown bear samples from different origins and different chronology, applying matrix-assisted laser desorption/ionisation time-of-flight mass spectrometry (MALDI TOF). We also performed an in-silico study of ursid bone collagen sequences published in databases. In our results we detected some discrepancies between the peptides obtained from both in silico and MALDI TOF analysis of fossil collagen and those published in the literature, in which we conclude that there are some misidentified peptides. The identification of skeletal remains by means of their peptide fingerprint is proving to be a powerful tool in palaeontology, which will bear greater fruit once the limitations of a technique that is in its initial stages have been overcome.

Byronids are problematic fossils of possible cnidarian affinity, only rarely reported from the Devonian, but never previously studied in the Barrandian area, Czech Republic. Two new species Prestephanoscyphus branzovensis sp. nov. and Prestephanoscyphus robustus sp. nov. are erected from the Lochkovian and the Eifelian, respectively. Four poorly known species referred to Byronia are described in open nomenclature from the Emsian and Eifelian. The new genus Parabyronia gen. nov. with the type species Parabyronia elegans sp. nov. is closely related to other sphenothallids but is distinguished by transverse ridges on its theca and short spines at the apertural end of the theca. Definite identification of phosphatic rings as the attachment structure of byronids was proved by new material of the Lochkovian age. A dwarf conularia Pidiconularia gen. nov. with the type species Pidiconularia tubulata sp. nov. is remarkable by its minute size and very fine ornament; its conulariid affinity is proved by subrectangular cross-section and four internal carinae. Microstructure of theca of Prestephanoscyphus is characterised by alternation of compact laminae of aligned columnar microcrystallites and chambered laminae with isometric microcrystallites of apatite. Accretionary growth of byronid theca and structure of the holdfast with basal opening for the adhesive pedal disc support their cnidarian affinity although they likely display the bilateral instead of tetramerous symmetry. The protective function of bilaterally symmetrical whorls of internal apophyses in Prestephanoscyphus is suggested and their role in strengthening of thecal wall or supporting function of gastric septa are disputed. The byronids are regarded as epibionts rather than attached to the rocks. They are representatives of benthic groups with phosphatic shells that declined with decreasing availability of phosphorus in seas and oceans.

The Lower Pleistocene Karnezeika locality, lies in the Peloponnese, southern Greece, and its fauna corresponds to the Middle Villafranchian biochronological unit (MN17). The recovered mammal assemblage includes, among others, a few remains of a large Cercopithecid. Herein, we describe this material, including an upper second molar, a partially preserved proximal radius and, possibly, an upper first incisor. The teeth show advanced stages of wear but retain their typical papionin characters, such as a strong lingual cleft and four bilophodont cusps in the molar. The general morphology and wear pattern of the teeth rules out the possibility that the remains belong to the genus Theropithecus, while the general size of the corresponding material excludes the possibility of a Macaca representative as well. On the contrary, the studied material better fits the size range of Paradolichopithecus. Even though this genus is likely represented in the Villafranchian of Europe by a single species, Par. arvernensis, the scarcity of the studied material imposes reservations and thus the Karnezeika papionin is referred at the moment to cf. Paradolichopithecus sp. As in the rest of Europe, the Paradolichopithecus record is rare in Greece, having been found in only two localities, Vatera and Dafnero. Despite its scarcity, the new material from Karnezeika indicates a wide distribution of this important taxon in the Greek peninsula.

A small lot of fossil whale barnacles from the Upper Pleistocene of California and the Middle Pleistocene (Chibanian) of Oregon (United States West Coast), described in a 1972 unpublished MA thesis, are formally described and illustrated herein. In that thesis, a new genus and species name were proposed; however, according to the International Code of Zoological Nomenclature, they have no taxonomic standing and are thus unavailable. Based on our reappraisal, two specimens in this lot belong to a new, extinct species that can be assigned to the purportedly extant genus Cetopirus. Cetopirus polysyrinx sp. nov. differs from congeners in that its secondary T-shaped flanges are multitubiferous internally, that is, they are perforated by a high number of irregularly-sized and irregularly-spaced tubules that result in a spongy aspect in transverse section. Whether or not this peculiar condition had any adaptive significance is difficult to determine. Considering that Cetopirus is currently known as an obligate epibiont of right whales (including the North Pacific form Eubalaena japonica (Lacépède 1818)), the host of C. polysyrinx sp. nov. was E. japonica or some other species of Eubalaena. The Plio-Pleistocene deposits of the Pacific coast of North America have yielded a rather idiosyncratic fossil whale barnacle fauna, inclusive of the genera Cetolepas, Cryptolepas and now Cetopirus, which seemingly contrasts with all other coeval assemblages worldwide, the latter being in turn dominated by Coronula spp.

Brachiopod taxonomy is based on descriptions of shell morphology and key characters, but diagenesis generally modifies or erases some of them, hindering brachiopod identification. Brachiopods that are taxonomically related usually present shells with similar appearance but can differ in size (i.e., Rhynchonellata). Some aspects of morphology – for example the angular measurement of the curvature of the shell or details of shell microstructure – could aid taxonomic identification. Gigantoproductids, which lack a robust taxonomy, have the largest shells among brachiopods and are ideal for this kind of study because of their gigantic size and morphological variability. Furthermore, they have a great abundance and worldwide distribution during the mid-Carboniferous. More than 700 samples have been collected from Sierra Morena (Spain), Montagne Noire (France) and Adarouch (Morocco) identifying up to six gigantoproductid genera: Globosoproductus, Semiplanus, Kansuella?, Latiproductus, Gigantoproductus and Datangia. Microstructural features from 170 thin sections belonging to gigantoproductid ventral valves have been studied, and six crystal morphologies have been distinguished within the tertiary layer: subhorizontal, imbricated, crenulated, acicular, short and long columnar morphologies. Moreover, 23 complete shells from all genera have been selected to investigate shell size and curvature. Results from this study emphasise that shell size, curvature and crystal shape are taxa-related. Finally, a remarkable morphological change in the gigantoproductid populations from the western Palaeo-Tethys occurred during the Viséan–Serpukhovian, from thin-shelled genera with subhorizontal morphology (Viséan) to thick-shelled genera with a tertiary layer consisting of long columnar crystals (Serpukhovian). This study proves that microstructure, maximum thickness and shell spiral characterisation are robust characters when applied to gigantoproductid taxonomy, but also have great potential in other brachiopod groups.

New 10Be exposure age dating and geomorphological mapping of emerged shoreline features in W Jura and NE Islay throw new light on the regional pattern of ice sheet deglaciation and late-glacial relative sea level change. We conclude that the oldest and highest emerged shorelines in this area were produced ~15.7–16.3 ka, shortly after ice sheet deglaciation ~16.5 ka. It is envisaged that the first incursion of marine waters into coastal areas took place close to a former ice sheet margin that oscillated in position across this part of the Scottish Inner Hebrides. The first evidence of late-glacial marine sedimentation following deglaciation consists of emerged marine terrace fragments and unvegetated gravel beach ridges, the former represented by a prominent glacio-isostatically tilted shoreline that declines in altitude NE to SW, from ~40 m above ordnance datum (OD) in NW Jura to ~19 m OD in central Islay. In W Jura, north of Loch Tarbert, spectacular staircases of up to 55 unvegetated gravel beach ridges were formed shortly after regional deglaciation, possibly within 1 ka. A preliminary estimate of the average rate of relative sea level lowering across W Jura between deglaciation and the Younger Dryas is in the order of ~7 mmyr−1. Geomorphological evidence from Shian Bay, W Jura, indicates a truncation of the late-glacial beach ridge staircases by a large 480-m-long beach ridge (the Colonsay Ridge) at ~14.9 ka, when former relative sea level was at ~18 m OD. This ridge may represent the product of either a stillstand in the progressive lowering of relative sea level during the late-glacial or a reversal. This raises the intriguing possibility of an association between ridge formation and the timing of the well-established global meltwater pulse 1A between ~14.65 and ~14.8 ka.

The geological and tectonic background of the Sabalan area in NW Iran and its present-day surface processes make it ideal for examining the effects of tectonic processes in shaping the Earth's crust. As a result of the intense distribution of pre-Quaternary and Quaternary structures (e.g., faults, joints and folds), most of the drainage basins in the southern and central parts of the study area have developed under tectonic-dominated conditions, whereas the effects of erosional processes are greater in the north and east. An evaluation of the geomorphic indices using the index of active tectonics (IAT) and analytical hierarchy process (AHP) methods shows that the AHP results are more reliable than the IAT results and are coherent with the geological and structural conditions of the study area. The geomorphic results are highly consistent with the intensity and distribution of fractures. The majority of fractures have developed in a NW–SE direction, indicating antithetic R′ Riedel fractures to the main NE-SW-trending faults. However, a significant number of the fractures in the study area are NNE–SSW- and NE–SW-oriented R and P fractures and NNW–SSE-oriented tension fractures. Palaeostress analysis of the fault data shows at least two faulting events in the pre-Quaternary and Quaternary, respectively. The pre-Quaternary NNW–SSE-striking dextral strike-slip faults experienced post-Eocene 25–30° clockwise rotation and re-activated as NE–SW-striking sinistral faults during the Quaternary. Although seismic activity is currently low, the consistency of our results with the regional stress data show that the study area is still tectonically active.

The presence of archaeocyath-bearing clasts from Cenozoic tills and Cambrian Mount Wegener Formation reveal erosion of a hidden Cambrian carbonate platform in Shackleton Range, Antarctica. We provide microfacies, paleontological, diagenetic and tectonically induced fabric data from carbonate clasts which, in addition to available geochemical and geochronological data from Shackleton Range, allow the paleoenvironmental reconstruction of a lost Cambrian Series 2 mixed siliciclastic–carbonate platform that was developed and eroded during the Ross orogeny. Carbonate production was dominated by non-skeletal grains in possibly restricted platform-interior and oolitic shoal complex settings, while open subtidal sub-environments (calcimicrobe carpets, calcimicrobe–archaeocyath patch reefs, muddy bottoms) were dominated by a diverse calcimicrobe assemblage and/or by secondary to accessory heterozoan assemblage (archaeocyaths and other sponges, chancelloriids, hyoliths, coralomorphs, trilobites, echinoderms). We describe a Botoman assemblage with 34 archaeocyathan species among 12 existing archaeocyathan genera. A new archaeocyath family Shackletoncyathidae is proposed. New species (Rotundocyathus glacius sp. nov., Buggischicyathus microporus gen. et sp. nov., Paragnaltacyathus hoeflei, Shackletoncyathus buggischi gen. et. sp. nov., Santelmocyathus santelmoi gen. et sp. nov., Wegenercyathus sexangulae gen. et sp. nov.) and Tabulaconus kordae coralomorph are reported from Antarctica for the first time. Archaeocyathan fauna share few species with contemporary fauna of South Australia (9) and even fewer with the Antarctic platforms of the Shackleton Limestone (2) or the Schneider Hills limestone (1). Similarity is greater with Antarctic allochthonous assemblages of Permo-Carboniferous tillites from Ellsworth Mountains (2), Cenozoic deposits from King George Island (4) or Weddell Sea (1). The Shackleton Range lost/hidden platform shows a distinct entity related with its tectonosedimentary evolution, in a possible back-arc basin on the Mozambique seaway during the E and W Gondwana amalgamation, which distinguishes it from those developed on the palaeo-Pacific margin of the E Antarctic craton.

The Jurassic to Cretaceous strata exposed in the Rollrock Section, Sverdrup Basin, Arctic Canada, represent one of the northernmost continuous outcrops spanning this poorly understood transition. The Oxfordian–Valanginian mudstones of the Rollrock Section were deposited in a shallow marine environment and, as such, provide the ideal lithology to investigate the response of high latitude dinoflagellate cyst populations to the frequent environmental perturbations of this time. Using a multivariate statistical approach, distinct palaeoecologically significant groups are identified and directly linked to time and palaeoenvironments, allowing for the reconstruction of underlying long term palaeoenvironmental trends. These palaeoenvironmental trends are identified to be driven by sequence stratigraphic cycles. For the first time, fourth order sequences are recorded from this pivotal period in the Sverdrup Basin and reveal an additional level of short term climatic events that complicate the identification of long term trends. The relationship existing between marine phytoplankton and climate is utilised to decipher the interplay of long-term and short-term climate fluctuations, distinguishing them from evolutionary trends. Two groups of dinoflagellate cysts, identified by process morphology, are recorded to act as environmental proxies. High percentages of proximochorate dinoflagellate cysts, such as Trichodinium erinaceoides, indicate more proximal, high energy, nutrient rich conditions and are dominant in fourth order transgressive systems tracts. High percentages of chorate dinoflagellate cysts, such as Oligosphaeridium complex, signify distal, low energy, nutrient depleted conditions and are dominant in fourth order highstand systems tracts.

The Paleogene lava flows of the Faroe Islands Basalt Group are divided into three relatively thick formations. The oldest, the Beinisvørð Formation is separated from the second lava flow succession, the Malinstindur Formation, by two formations composed primarily of volcaniclastic rocks. The oldest of these, the Prestfjall Formation has been interpreted as a period of eruptive quiescence and linked to changes in mantle melting. It is characterised in the south by the occurrence of coals, while the overlying Hvannhagi Formation is a sequence of primary and remobilised volcaniclastic strata. Field, laboratory, palynology, and photogrammetry studies have been used to investigate variations in facies and architecture within these volcaniclastic formations. The data reveal significantly different depositional systems in the Prestfjall and Hvannhagi formations over the ~40 km from the island of Vágar in the north to the island of Suðuroy in the south. Facies distribution in both the Prestfjall and Hvannhagi formations was found to have been controlled by a complex interaction of regional paleoslope, pre-existing topography, the eruption and local collapse of low-angle shield volcanoes, and minor brittle deformation. Lithological data and photogrammetry have enabled the identification of a > 180 m thick succession of volcaniclastic conglomerates deposited by lahars reworking a low-angle shield sector collapse. Co-occurrence of facies characteristic of the Prestfjall, Hvannhagi and Malinstindur formations indicate that volcanic eruption continued at a lower tempo throughout the Prestfjall Formation interval. Identification of a Beinisvørð Formation low-angle volcano shield northwest of the Faroe Islands alters the previous eruption model for this extensive lava field.

Source materials for investigating the life of James Croll are examined and collated. This is organised around the topics of: Croll's Autobiographical sketch and the Memoir of his life and work, both contained within the volume produced by James Campbell Irons; publications by Croll; aspects of his genealogy; manuscript sources in publicly accessible archives and in private ownership; and other published sources.

Popular and scholarly information concerning the life of James Croll has been accumulating slowly since the death in 1890 of the self-taught climate change pioneer. The papers in the current volume offer thorough assessments of topics associated with Croll's work, but this contribution seeks to provide a personal context for an understanding of James Croll the man, as well as James Croll the scholar of sciences and religion. Using archival as well as published sources, emphasis is placed upon selected components of his life and some of the less recognised features of his biography. These include his family history, his many homes, his health, participation in learned societies and attitudes to collegiality, financial problems including the failed efforts to secure a larger pension, and friendship. Life delivered a mixture of ‘trials and sorrows’, but it seems clear from the affection and respect accorded him that many looked upon James Croll as a ‘man greater far than his work’.

The integration of biostratigraphical, wireline log, geophysical and available geochronological ages has identified two principal periods of volcanism in the Faroe–Shetland and Rockall basins. The first is pre-breakup, upper Danian to lower Thanetian: in the Rockall and Faroe–Shetland basins, isolated volcanic activity from 62 Ma to 58.7 Ma is identified in areas closely linked to the SSW–NNE structural fabric of the continental margin. Volcanic activity was concentrated at basin flank fissures and localised point sources. This rift-flank volcanism led to widespread volcanic ash deposition, localised lava flow fields and the formation of igneous centres. Some of the Hebridean and onshore central complexes (e.g., Rum) were uplifted and rapidly eroded during the later pre-breakup period, while additional accommodation space was developed in the adjacent offshore basins. Onset and termination of pre-breakup volcanism is correlated to intra-plate stress regimes in Europe, following the cessation of convergence of Africa and Europe in the Danian. The second is syn-breakup, upper Thanetian to Ypresian, initiated at ca.57 Ma in the Rockall and Faroe–Shetland basins. Initial high-volume extrusive igneous successions were focussed to the W in the Faroe–Shetland Basin. In the centre and E of the Faroe–Shetland and Rockall basins, separate eruption loci developed along pre-existing lineaments either as fissure or point-sourced lava fields. Short-term cessation of eruption at ~55.8 Ma was followed by resumption of flood basalt eruptions and a shift in focus to the NW. Fluctuations in the syn-breakup eruption tempo are reflected in the formation and subsequent rejuvenation of prominent unconformities, only previously recognised as a single erosive event. The W and northward shift of eruption focus, and the eruption of mid ocean ridge basalt-type lavas in the syn-breakup period reflect the onset of lithospheric thinning in the nascent North Atlantic Rift prior to flooding of the rift and eruption of the widespread lower Ypresian Balder Formation tephras.

James Croll left school at the age of 13 years, yet while a janitor in Glasgow he published a landmark paper on astronomically-related climate change, claimed as ‘the most important discovery in paleoclimatology’, and which brought him to the attention of Charles Darwin, William Thomson and John Tyndall, amongst others. By 1867 he was persuaded to become Secretary and Accountant of the newly established Geological Survey of Scotland in Edinburgh, and a year after the appearance of his keynote volume Climate and time in 1875, he was lauded with an honorary doctorate from Scotland's oldest university, Fellowship of the Royal Society of London and Honorary Membership of the New York Academy of Sciences. Using a range of archival and published sources, this paper explores aspects of his ‘journey’ and the background to the award of these major accolades. It also discusses why he never became a Fellow of his national academy, the Royal Society of Edinburgh. In the world of 19th-Century science, Croll was not unusual in being both an autodidact and of humble origins, nor was he lacking in support for his endeavours. It is possible that a combination of Croll's modesty and innovative genius fostered advancement, though this did not hinder a willingness to engage in vigorous argument.

James Croll was a pioneer in studies of the impact of the slowly changing orbital dynamics of the Earth on climate change. His book Climate and Time in their Geological Relations (1875) was far ahead of its time in seeking correlations between climate change, the occurrence of ice ages and perturbations to the Earth's orbit about the Sun. The astronomical cycles he discovered are now called ‘Milankovitch Cycles’ after the Serbian scientist whose research was first published in the Handbuch der Klimatologie in 1930. The celestial mechanical and astronomical background to Croll's research is the focus of this essay. The development of the understanding of the impact of perturbations of the elliptical planetary orbits by other bodies in the solar system paralleled new mathematical techniques, many of which were developed in association with celestial mechanical problems. The central contributions of many of the major mathematicians of the late 18th and 19th Centuries, including Euler, Lagrange, Laplace and Le Verrier, are highlighted. Although Croll's contributions faded from view for several generations, his pioneering insights have now been demonstrated to have been basically correct.

Science, for James Croll, began and ended in metaphysics. Metaphysics, in turn, provided proof of a First and Final Cause of all things. This proof rested on two metaphysical principles: that every event must have a cause, and that the determination of a cause is distinct from its production. This argument emerged from his deeply held religious commitments. As a 17-year-old, he converted to a Calvinist and evangelical form of Christianity. After a period of questioning the Calvinist system, he embraced it again through reading the famous treatise on the will by the New England theologian, Jonathan Edwards. This determinedly metaphysical work, which engaged as much with Enlightenment thought as with Calvinism, defended the view that the will was not a self-determining cause of human action. This ‘hard case’ provided the basis for a larger claim that every act whatever has a cause, and that the production of an act was different from its determination. In part through reading Edwards, Croll remained a devout and convinced ‘moderate’ Calvinist for the rest of his life. He also developed a deep love of metaphysics and became convinced that without it, everything, including science, remained confused and in darkness. For Croll, even the most basic science could not be properly conducted without prior metaphysical principles. But this was more than just an argument about the philosophical foundations of scientific inquiry. It was also based on Croll's conviction that the cosmos, earth history and life (including his own) was fully determined by a supreme and perfect intellect. This conviction entered into the marrow of Croll's scientific theories and shaped his interpretation of the twists and turns of his own life. In short, to take seriously Croll's own self-understanding, we need to allow him to ‘do God’.

Scientific societies played a crucial role in the emergence of a professional culture of science in Britain in the mid- to late-19th Century. At first sight, James Croll's membership of a limited number of scientific associations may be assumed to be the result of his lack of social credit and scientific connections. In this article, by examining Croll's correspondence, I demonstrate that Croll's select participation in scientific clubs and associations reflected his strategic pursuit of a vision of science set apart from party or societal affiliation. I focus on the contrasting histories of the Royal Society of Edinburgh and the Geological Survey, as well as the institutional history of the Philosophical Magazine. Situating the institutions in their respective social and cultural contexts, I argue that the more meritocratic, inclusive social structure of the Survey and Magazine helps explain Croll's choice to avoid affiliation with the Royal Society of Edinburgh.