Agnew, W., Uresk, D. W., & Hansen, R. M. (1986) Flora and fauna associated with prairie dog colonies and adjacent ungrazed mixed-grass prairie in western South Dakota. Journal of Range Management 39: 135–139.
Anderson, T. M., Fokkema, W., Valls-Fox, H., & Olff, H. (2013) Distinct physiological responses underlie defoliation tolerance in African lawn and bunch grasses. International Journal of Plant Sciences 174: 769–778.
Archibald, S. (2008) African grazing lawns – how fire, rainfall, and grazer numbers interact to affect grass community states. The Journal of Wildlife Management 72: 492–501.
Archibald, S. & Bond, W. J. (2004) Grazer movements: spatial and temporal responses to burning in a tall-grass African savanna. International Journal of Wildland Fire 13: 1–9.
Archibald, S., Bond, W. J., Stock, W. D., & Fairbanks, D. H. K. (2005) Shaping the landscape: fire–grazer interactions in an African savanna. Ecological Applications 15: 96–109.
Arnold, S. G., Anderson, T. M., & Holdo, R. M. (2014) Edaphic, nutritive, and species assemblage differences between hotspots and matrix vegetation: two African case studies. Biotropica 46: 387–394.
Arsenault, R. & Owen-Smith, N. (2008) Resource partitioning by grass height among grazing ungulates does not follow body size relation. Oikos 117: 1711–1717.
Arsenault, R. & Owen-Smith, N. (2011) Competition and coexistence among short-grass grazers in the Hluhluwe-iMfolozi Park, South Africa. Canadian Journal of Zoology 89: 900–907.
Arshad, M. A. (1982) Influence of the termite Macrotermes michaelseni (Sjost) on soil fertility and vegetation in a semi-arid savannah ecosystem. Agro-Ecosystems 8: 47–58.
Askins, R. A., Chávez-Ramírez, F., Dale, B. C., et al. (2007) Conservation of grassland birds in North America: understanding ecological processes in different regions. Ornithological Monographs 64: 1–46.
Augustine, D. J. (2003) Long-term, livestock-mediated redistribution of nitrogen and phosphorus in an East African savanna. Journal of Applied Ecology 40: 137–149.
Augustine, D. J. & McNaughton, S. J. (1998) Ungulate effects on the functional species composition of plant communities: herbivore selectivity and plant tolerance. Journal of Wildlife Management 62: 1165–1183.
Augustine, D. J. & McNaughton, S. J. (2006) Interactive effects of ungulate herbivores, soil fertility, and variable rainfall on ecosystem processes in a semi-arid savanna. Ecosystems 9: 1242–1256.
Augustine, D. J. & Skagen, S. K. (2014) Mountain plover nest survival in relation to prairie dog and fire dynamics in shortgrass steppe. Journal of Wildlife Management 78: 595–602.
Augustine, D. J., McNaughton, S. J., & Frank, D. A. (2003) Feedbacks between soil nutrients and large herbivores in a managed savanna ecosystem. Ecological Applications 13: 1325–1337.
Bartlett, M. K., Scoffoni, C., & Sack, L. (2012) The determinants of leaf turgor loss point and prediction of drought tolerance of species and biomes: a global meta-analysis. Ecology Letters 15: 393–405.
Bell, R. H. V. (1971) A grazing ecosystem in the Serengeti. Scientific American 225: 86–94.
Blackmore, A. C., Mentis, M. T., & Scholes, R. J. (1990) The origin and extent of nutrient-enriched patches within a nutrient-poor savanna in South Africa. Journal of Biogeography 17: 463–470.
Bond, W. J., Smythe, K., & Balfour, D. A. (2001) Acacia species turnover in space and time in an African savanna. Journal of Biogeography 28: 117–128.
Bonnet, O., Fritz, H., Gignoux, J., & Meuret, M. (2010) Challenges of foraging on a high-quality but unpredictable food source: the dynamics of grass production and consumption in savanna grazing lawns. Journal of Ecology 98: 908–916.
Brooks, P. M. & Macdonald, I. A. W. (1983) The Hluhluwe-Umfolozi Reserve: an ecological case history. In: Management of large mammals in African conservation areas (ed. Owen-Smith, N.), pp. 51–77. Haum Educational Publishers, Pretoria.
Brooks, S. J. (2001) Changing nature: a critical historical geography of the Umfolozi and Hluhluwe Game Reserves, Zululand, 1887 to 1947. PhD thesis, Queen's University, Kingston, Canada.
Clauss, M., Castell, J. C., Kienzle, E., et al. (2007) Mineral absorption in the black rhinoceros (Diceros bicornis) as compared with the domestic horse. Journal of Animal Physiology and Animal Nutrition 91: 193–204.
Coetsee, C., Stock, W. D., & Craine, J. M. (2010) Do grazers alter nitrogen dynamics on grazing lawns in a South African savannah? African Journal of Ecology 49: 62–69.
Coughenour, M. B. (1985) Graminoid responses to grazing by large herbivores – adaptations, exaptations, and interacting processes. Annals of the Missouri Botanical Garden 72: 852–863.
Craig, T. P. (2010) The resource regulation hypothesis and positive feedback loops in plant–herbivore interactions. Population Ecology 52: 461–473.
Cromsigt, J. P. G. M. (2006) Large herbivores in space: resource partitioning among savanna grazers in a heterogeneous environment. PhD thesis, University of Groningen, Groningen.
Cromsigt, J. P. G. M. & Kuijper, D. P. J. (2011) Revisiting the browsing lawn concept: evolutionary interactions or pruning herbivores? Perspectives in Plant Ecology, Evolution or Systematics 13: 207–215.
Cromsigt, J. P. G. M. & Olff, H. (2008) Dynamics of grazing lawn formation: an experimental test of the role of scale-dependent processes. Oikos 117: 1444–1452.
Cromsigt, J. P. G. M. & te Beest, M. (2014) Restoration of a megaherbivore: landscape-level impacts of white rhinoceros in Kruger National Park, South Africa. Journal of Ecology 102: 566–575.
Cromsigt, J. P. G. M., Prins, H. H. T. & Olff, H. (2009) Habitat heterogeneity as a driver of ungulate diversity and distribution patterns: interaction of body mass and digestive strategy. Diversity and Distributions 15: 513–522.
Currie, G. (2003) The impact of megaherbivore grazers on grasshopper communities via grassland conversion in a savannah ecosystem. Honours thesis, University of Cape Town, Cape Town.
Davies, A. B., Robertson, M. P., Levick, S. R., et al. (2014) Variable effects of termite mounds on African savanna grass communities across a rainfall gradient. Journal of Vegetation Science 25: 1405–1416.
Deane, N. N. (1966) Ecological changes and their effect on a population of reedbuck (Redunca arundinum (Boddaert)). Lammergeyer 6: 2–8.
Desmond, M. J. (2004) Effects of grazing practices and fossorial rodents on a winter avian community in Chihuahua, Mexico. Biological Conservation 116: 235–242.
Downing, B. H. (1972) A plant ecological survey of the Imfolozi Game Reserve, Zululand. PhD thesis, University of Natal, Durban.
Du Plessis, S. S. (1972) Ecology of blesbok with special reference to productivity. Wildlife Monographs 30: 1–70.
Eltringham, S. K. (1974) Changes in the large mammal community of Mweya Peninsula, Rwenzori National Park, Uganda, following removal of hippopotamus. Journal of Applied Ecology 11: 855–865.
Feely, J. M. (1980) Did Iron Age man have a role in the history of Zululand's wilderness landscapes? South African Journal of Science 76: 150–152.
Feely, J. M. (2004) Prehistoric use of woodland and forest by farming peoples in South Africa. In: Indigenous forests and woodlands: policy, people and practice (eds Lawes, M. J., Eeley, H. A. C., Shackleton, C. M., & Geach, B. G. S.), pp. 284–286. University of KwaZulu-Natal Press, Pietermaritzburg.
Frank, D. A., Groffman, P. M., Evans, R. D., & Tracy, B. F. (2000) Ungulate stimulation of nitrogen cycling and retention in Yellowstone Park grasslands. Oecologia 123: 116–121.
Fuhlendorf, S. D. & Engle, D. M. (2001) Restoring heterogeneity on rangelands: ecosystem management based on evolutionary grazing patterns. BioScience 51: 625–632.
Gandar, M. V., Huntley, B. J., & Walker, B. H. (1982) Trophic ecology and plant/herbivore energetics. In: Ecology of tropical savannas (eds Huntley, B. J. & Walker, B. H.), pp. 514–534. Springer Verlag, Berlin.
Girma, F. S. & Krieg, D. R. (1992) Osmotic adjustment in sorghum. 1. Mechanisms of diurnal osmotic potential changes. Plant Physiology 99: 577–582.
Gosling, C. M. (2014) Biotic determinants of heterogeneity in a South African savanna. PhD thesis, University of Groningen, Groningen.
Gosling, C. M., Cromsigt, J. P. G. M., & Olff, H. (2011) Effects of erosion from mounds of different termite genera on distinct functional grassland types in an African savanna. Ecosystems 15: 128–139.
Grant, C. C. & Scholes, M. C. (2006) The importance of nutrient hot-spots in the conservation and management of large wild mammalian herbivores in semi-arid savannas. Biological Conservation 130: 426–437.
Hagenah, N., Prins, H. H. T., & Olff, H. (2009) Effects of large herbivores on murid rodents in a South African savanna. Journal of Tropical Ecology 25: 483–492.
Hall, M. (1984) Prehistoric farming in the Mfolozi and Hluhluwe valleys of southeast Africa: an archaeo-botanical survey. Journal of Archaeological Science 11: 223–235.
Hempson, G. P., Archibald, S., Bond, W. J., et al. (2015) Ecology of grazing lawns in Africa. Biological Reviews 90: 979–994.
Illius, A. W. & O'Connor, T. G. (2000) Resource heterogeneity and ungulate population dynamics. Oikos 89: 283–294.
Jouquet, P., Boulain, N., Gignoux, J. & Lepage, M. (2004) Association between subterranean termites and grasses in a West African savanna: spatial pattern analysis shows a significant role for Odontotermes n. pauperans. Applied Soil Ecology 27: 99–107.
Karki, J. B., Jhala, Y. V., & Khanna, P. P. (2000) Grazing lawns in Terai Grasslands, Royal Bardia National Park, Nepal. Biotropica 32: 423–429.
Kim, H., Anderson, S. H., Motavalli, P. P., & Gantzer, C. J. (2010) Compaction effects on soil macropore geometry and related parameters for an arable field. Geoderma 160: 244–251.
Kleynhans, E. J., Jolles, A. E., Bos, M. R. E., & Olff, H. (2011) Resource partitioning along multiple niche dimensions in differently sized African savanna grazers. Oikos 120: 591–600.
Knapp, A. K., Blair, J. M., Briggs, J. M., et al. (1999) The keystone role of bison in North American tallgrass prairie. BioScience 49: 39–50.
Krook, K., Bond, W. J., & Hockey, P. A. R. (2007) The effect of grassland shifts on the avifauna of a South African savanna. Ostrich – Journal of African Ornithology 78: 271–279.
Lock, J. M. (1972) The effects of hippopotamus grazing on grassland. Journal of Ecology 60: 445–467.
McNaughton, S. J. (1983) Compensatory growth as a response to herbivory. Oikos 40: 329–336.
McNaughton, S. J. (1984) Grazing lawns: animals in herds, plant form, and coevolution. American Naturalist 124: 863–886.
McNaughton, S. J. (1985) Ecology of a grazing ecosystem: the Serengeti. Ecological Monographs 55: 259–294.
McNaughton, S. J. (1988) Mineral-nutrition and spatial concentrations of African ungulates. Nature 334: 343–345.
McNaughton, S. J., Banyikwa, F. F., & McNaughton, M. M. (1997) Promotion of the cycling of diet-enhancing nutrients by African grazers. Science 278(5344): 1798–1800.
Mgobozi, M. P. (2008) Spider community responses to Chromolaena odorata invasion, grassland type and grazing intensities. MSc thesis, University of Pretoria, Pretoria.
Milchunas, D. G., Sala, O. E., & Lauenroth, W. K. (1988) A generalized-model of the effects of grazing by large herbivores on grassland community structure. American Naturalist 132: 87–106.
Mills, A. J., Milewski, A., Fey, M. V., Groengroeft, A., & Petersen, A. (2009) Fungus culturing, nutrient mining and geophagy: a geochemical investigation of Macrotermes and Trinervitermes mounds in southern Africa. Journal of Zoology 278: 24–35.
Morgan, J. W. & Lunt, I. D. (1999) Effects of time-since-fire on the tussock dynamics of a dominant grass (Themeda triandra) in a temperate Australian grassland. Biological Conservation 88: 379–386.
Novellie, P. (1990) Habitat use by indigenous grazing ungulates in relation to sward structure and veld condition. Journal of the Grassland Society of Southern Africa 7: 16–23.
Novellie, P. & Gaillard, A. (2013) Long-term stability of grazing lawns in a small protected area, the Mountain Zebra National Park. Koedoe 55: 1–7.
Olivier, R. C. D. & Laurie, W. A. (1974) Habitat utilization by hippopotamus in the Mara river. African Journal of Ecology 12: 249–271.
Owen-Smith, N. (1973) The behavioural ecology of the white rhinoceros. PhD thesis, University of Wisconsin, Madison, WI.
Owen-Smith, N. (1988) Megaherbivores: the influence of very large body size on ecology. Cambridge University Press, Cambridge.
Owen-Smith, N. (2004) Functional heterogeneity in resources within landscapes and herbivore population dynamics. Landscape Ecology 19: 761–771.
Person, B. T., Herzog, M. P., Ruess, R. W., et al. (2003) Feedback dynamics of grazing lawns: coupling vegetation change with animal growth. Oecologia 135: 583–592.
Roberts, C. M. (2009) Marsupial grazing lawns in Tasmania: maintenance, biota and the effects of climate change. PhD thesis, University of Tasmania.
Scholes, R. J. (2003) Convex relationships in ecosystems containing mixtures of trees and grass. Environmental and Resource Economics 26: 559–574.
Shrader, A. M. & Perrin, M. R. (2006) Influence of density on the seasonal utilization of broad grassland types by white rhinoceroses. African Zoology 41: 312–315.
Stock, W. D., Bond, W. J., & Van de Vijver, C. A. D. M. (2010) Herbivore and nutrient control of lawn and bunch grass distributions in a southern African savanna. Plant Ecology 206: 15–27.
Swemmer, T. (1998) The distribution and ecology of grazing lawns in a South African savannah ecosystem. Honours thesis, University of Cape Town, Cape Town.
Tracy, B. & McNaughton, S. J. (1995) Elemental analysis of mineral lick soils from the Serengeti National Park, the Konza Prairie and Yellowstone National Park. Ecography 18: 91–94.
Valls-Fox, H., Bonnet, O., Cromsigt, J. P. G. M., Fritz, H., & Shrader, A. M. (2015) Legacy effects of different land-use histories interact with current grazing patterns to determine grazing lawn properties. Ecosystems 18: 720–733.
Van de Vijver, C. A. D. M., Poot, P., & Prins, H. H. T. (1999) Causes of increased nutrient concentrations in post-fire regrowth in an East African savanna. Plant and Soil 214: 173–185.
Van der Plas, F., Anderson, T. M., & Olff, H. (2012) Trait similarity patterns within grass and grasshopper communities: multitrophic community assembly at work. Ecology 93: 836–846.
Van der Plas, F., Zeinstra, P., Veldhuis, M., et al. (2013) Responses of savanna lawn and bunch grasses to water limitation. Plant Ecology 214: 1157–1168.
Van der Waal, C., Kool, A., Meijer, S. S., et al. (2011) Large herbivores may alter vegetation structure of semi-arid savannas through soil nutrient mediation. Oecologia 165: 1095–1107.
Veldhuis, M. P., Howison, R. A., Fokkema, R. W., Tielens, E., & Olff, H. (2014) A novel mechanism for grazing lawn formation: large herbivore-induced modification of the plant–soil water balance. Journal of Ecology 102: 1506–1517.
Veldhuis, M. P., Fakkert, H. F., Berg, M. P., & Olff, H. (2016). Grassland structural heterogeneity in a savanna is driven more by productivity differences than by consumption differences between lawn and bunch grasses. Oecologia 182: 841–853.
Verweij, R. J. T., Verrelst, J., Loth, P. E., Heitkönig, I. M. A. & Brunsting, A. M. H. (2006) Grazing lawns contribute to the subsistence of mesoherbivores on dystrophic savannas. Oikos 114: 108–116.
Vesey-Fitzgerald, D. F. (1960) Grazing succession among East African game animals. Journal of Mammalogy 41: 161–172.
Vos, I. A. (2010) Spatial heterogeneity of resources – variation in space and time in the distribution and persistence of grazing lawns. MSc thesis, Utrecht University, Utrecht.
Waldram, M. S., Bond, W.J., & Stock, W. D. (2008) Ecological engineering by a mega-grazer: white rhino impacts on a South African savanna. Ecosystems 11: 101–112.
Walters, M., Midgley, J. J., & Somers, M. J. (2004) Effects of fire and fire intensity on the germination and establishment of Acacia karroo, Acacia nilotica, Acacia luederitzii and Dichrostachys cinerea in the field. BMC Ecology 4: 1–13.
Wood, T. G. & Sands, W. (1978) The role of termites in ecosystems. In: Production ecology of ants and termites (ed. Brian, M. V.), pp. 245–292. Cambridge University Press, Cambridge.
Young, T. P., Patridge, N., & Macrae, A. (1995) Long-term glades in acacia bushland and their edge effects in Laikipia, Kenya. Ecological Applications 5: 97–108.