%PDF-1.3
%
1 0 obj
<<
/Names 2 0 R
/Outlines 3 0 R
/Metadata 4 0 R
/Pages 5 0 R
/Type /Catalog
/PageLabels 6 0 R
>>
endobj
7 0 obj
<<
/CreationDate (D:20201008213402+05'30')
/Creator (Arbortext Advanced Print Publisher 10.0.1465/W Unicode)
/Producer (Acrobat Distiller 8.1.0 \(Windows\))
/ModDate (D:20220416093935+00'00')
/Title (S1431927620024320jra 978..988)
>>
endobj
2 0 obj
<<
/Dests 8 0 R
>>
endobj
3 0 obj
<<
/First 9 0 R
/Count 1
/Last 9 0 R
>>
endobj
4 0 obj
<<
/Subtype /XML
/Length 1732
/Type /Metadata
>>
stream
application/pdf
S1431927620024320jra 978..988
Not for further distribution unless allowed by the License or with the express written permission of Cambridge University Press.
https://doi.org/10.1017/S1431927620024320
https://doi.org/10.1017/S1431927620024320 Published online by Cambridge University Press
2020-10-08T21:34:02+05:01
Arbortext Advanced Print Publisher 10.0.1465/W Unicode
2022-04-16T09:39:35+00:00
Acrobat Distiller 8.1.0 (Windows)
uuid:809b1e02-b4de-4495-bdb4-6a8acc015f24
uuid:1f8d9ce6-c18c-4f2a-88e6-4e87aad46a6e
endstream
endobj
5 0 obj
<<
/Count 11
/Type /Pages
/Kids [10 0 R 11 0 R]
>>
endobj
6 0 obj
<<
/Nums [0 12 0 R]
>>
endobj
8 0 obj
<<
/Kids [13 0 R 14 0 R]
>>
endobj
9 0 obj
<<
/First 15 0 R
/Parent 3 0 R
/Dest
/Count -6
/Last 16 0 R
/Title
>>
endobj
10 0 obj
<<
/Parent 5 0 R
/Count 10
/Type /Pages
/Kids [17 0 R 18 0 R 19 0 R 20 0 R 21 0 R 22 0 R 23 0 R 24 0 R 25 0 R 26 0 R]
>>
endobj
11 0 obj
<<
/Parent 5 0 R
/Count 1
/Type /Pages
/Kids [27 0 R]
>>
endobj
12 0 obj
<<
/St 978
/S /D
>>
endobj
13 0 obj
<<
/Limits [ ]
/Names [ 28 0 R 29 0 R 30 0 R 31 0 R 32 0 R
33 0 R 34 0 R 35 0 R 36 0 R 37 0 R
38 0 R 39 0 R 40 0 R 41 0 R 42 0 R
43 0 R 44 0 R 45 0 R 46 0 R 47 0 R
48 0 R 49 0 R 50 0 R 51 0 R 52 0 R
53 0 R 54 0 R 55 0 R 56 0 R 57 0 R
58 0 R 59 0 R 60 0 R 61 0 R 62 0 R
63 0 R 64 0 R 65 0 R 66 0 R 67 0 R
68 0 R 69 0 R 70 0 R 71 0 R 72 0 R
73 0 R 74 0 R 75 0 R 76 0 R 77 0 R
78 0 R 79 0 R 80 0 R 81 0 R 82 0 R
83 0 R 84 0 R 85 0 R 86 0 R 87 0 R
88 0 R 89 0 R 90 0 R 91 0 R]
>>
endobj
14 0 obj
<<
/Limits [ ]
/Names [ 92 0 R 93 0 R 94 0 R 95 0 R 96 0 R
97 0 R 98 0 R 99 0 R 100 0 R 101 0 R
102 0 R 103 0 R 104 0 R 105 0 R 106 0 R
107 0 R 108 0 R 109 0 R]
>>
endobj
15 0 obj
<<
/Parent 9 0 R
/Next 110 0 R
/Dest
/Title
>>
endobj
16 0 obj
<<
/Parent 9 0 R
/Dest
/Prev 111 0 R
/Title
>>
endobj
17 0 obj
<<
/CropBox [0 0 593.972 792]
/Annots [112 0 R 113 0 R 114 0 R 115 0 R 116 0 R 117 0 R 118 0 R 119 0 R 120 0 R 121 0 R
122 0 R 123 0 R 124 0 R 125 0 R 126 0 R 127 0 R 128 0 R 129 0 R 130 0 R 131 0 R
132 0 R 133 0 R 134 0 R 135 0 R 136 0 R 137 0 R 138 0 R 139 0 R 140 0 R 141 0 R
142 0 R]
/Parent 10 0 R
/Contents [143 0 R 144 0 R 145 0 R 146 0 R 147 0 R 148 0 R 149 0 R 150 0 R 151 0 R 152 0 R
153 0 R 154 0 R]
/Rotate 0
/MediaBox [0 0 593.972 792]
/Resources 155 0 R
/Type /Page
>>
endobj
18 0 obj
<<
/CropBox [0 0 593.972 792]
/Annots [156 0 R 157 0 R 158 0 R 159 0 R 160 0 R 161 0 R 162 0 R 163 0 R 164 0 R 165 0 R
166 0 R 167 0 R]
/Parent 10 0 R
/Contents [168 0 R 169 0 R 170 0 R]
/Rotate 0
/MediaBox [0 0 593.972 792]
/Resources 171 0 R
/Type /Page
>>
endobj
19 0 obj
<<
/CropBox [0 0 593.972 792]
/Annots [172 0 R 173 0 R 174 0 R 175 0 R 176 0 R 177 0 R 178 0 R 179 0 R 180 0 R]
/Parent 10 0 R
/Contents [181 0 R 182 0 R 183 0 R]
/Rotate 0
/MediaBox [0 0 593.972 792]
/Resources 184 0 R
/Type /Page
>>
endobj
20 0 obj
<<
/CropBox [0 0 593.972 792]
/Annots [185 0 R 186 0 R 187 0 R 188 0 R 189 0 R 190 0 R 191 0 R 192 0 R 193 0 R 194 0 R
195 0 R 196 0 R 197 0 R 198 0 R]
/Parent 10 0 R
/Contents [199 0 R 200 0 R 201 0 R]
/Rotate 0
/MediaBox [0 0 593.972 792]
/Resources 202 0 R
/Type /Page
>>
endobj
21 0 obj
<<
/CropBox [0 0 593.972 792]
/Annots [203 0 R 204 0 R 205 0 R 206 0 R]
/Parent 10 0 R
/Contents [207 0 R 208 0 R 209 0 R]
/Rotate 0
/MediaBox [0 0 593.972 792]
/Resources 210 0 R
/Type /Page
>>
endobj
22 0 obj
<<
/CropBox [0 0 593.972 792]
/Annots [211 0 R 212 0 R 213 0 R 214 0 R 215 0 R 216 0 R 217 0 R 218 0 R 219 0 R 220 0 R
221 0 R 222 0 R]
/Parent 10 0 R
/Contents [223 0 R 224 0 R 225 0 R]
/Rotate 0
/MediaBox [0 0 593.972 792]
/Resources 226 0 R
/Type /Page
>>
endobj
23 0 obj
<<
/CropBox [0 0 593.972 792]
/Annots [227 0 R 228 0 R 229 0 R 230 0 R 231 0 R 232 0 R 233 0 R 234 0 R]
/Parent 10 0 R
/Contents [235 0 R 236 0 R 237 0 R]
/Rotate 0
/MediaBox [0 0 593.972 792]
/Resources 238 0 R
/Type /Page
>>
endobj
24 0 obj
<<
/CropBox [0 0 593.972 792]
/Annots [239 0 R 240 0 R 241 0 R 242 0 R]
/Parent 10 0 R
/Contents [243 0 R 244 0 R 245 0 R]
/Rotate 0
/MediaBox [0 0 593.972 792]
/Resources 246 0 R
/Type /Page
>>
endobj
25 0 obj
<<
/CropBox [0 0 593.972 792]
/Parent 10 0 R
/Contents [247 0 R 248 0 R 249 0 R]
/Rotate 0
/MediaBox [0 0 593.972 792]
/Resources 250 0 R
/Type /Page
/Annots [251 0 R]
>>
endobj
26 0 obj
<<
/CropBox [0 0 593.972 792]
/Annots [252 0 R 253 0 R 254 0 R 255 0 R 256 0 R]
/Parent 10 0 R
/Contents [257 0 R 258 0 R 259 0 R]
/Rotate 0
/MediaBox [0 0 593.972 792]
/Resources 260 0 R
/Type /Page
>>
endobj
27 0 obj
<<
/CropBox [0 0 593.972 792]
/Annots [261 0 R 262 0 R 263 0 R 264 0 R]
/Parent 11 0 R
/Contents [265 0 R 266 0 R 267 0 R]
/Rotate 0
/MediaBox [0 0 593.972 792]
/Resources 268 0 R
/Type /Page
>>
endobj
28 0 obj
<<
/D [19 0 R /FitR 30 662 297 634]
>>
endobj
29 0 obj
<<
/D [18 0 R /FitR 298 734 565 509]
>>
endobj
30 0 obj
<<
/D [18 0 R /FitR 298 333 565 40]
>>
endobj
31 0 obj
<<
/D [20 0 R /FitR 298 734 565 126]
>>
endobj
32 0 obj
<<
/D [21 0 R /FitR 30 734 297 537]
>>
endobj
33 0 obj
<<
/D [25 0 R /FitR 30 734 297 169]
>>
endobj
34 0 obj
<<
/D [25 0 R /FitR 294 734 560 527]
>>
endobj
35 0 obj
<<
/D [25 0 R /FitR 294 419 560 39]
>>
endobj
36 0 obj
<<
/D [26 0 R /FitR 35 734 301 300]
>>
endobj
37 0 obj
<<
/D [26 0 R /FitR 298 537 565 510]
>>
endobj
38 0 obj
<<
/D [26 0 R /FitR 298 317 565 290]
>>
endobj
39 0 obj
<<
/D [26 0 R /FitR 298 297 565 270]
>>
endobj
40 0 obj
<<
/D [26 0 R /FitR 298 287 565 260]
>>
endobj
41 0 obj
<<
/D [26 0 R /FitR 298 257 565 230]
>>
endobj
42 0 obj
<<
/D [26 0 R /FitR 298 227 565 200]
>>
endobj
43 0 obj
<<
/D [26 0 R /FitR 298 197 565 170]
>>
endobj
44 0 obj
<<
/D [26 0 R /FitR 298 167 565 140]
>>
endobj
45 0 obj
<<
/D [26 0 R /FitR 298 147 565 120]
>>
endobj
46 0 obj
<<
/D [26 0 R /FitR 298 127 565 100]
>>
endobj
47 0 obj
<<
/D [26 0 R /FitR 298 97 565 70]
>>
endobj
48 0 obj
<<
/D [26 0 R /FitR 298 517 565 490]
>>
endobj
49 0 obj
<<
/D [26 0 R /FitR 298 77 565 50]
>>
endobj
50 0 obj
<<
/D [27 0 R /FitR 30 739 297 712]
>>
endobj
51 0 obj
<<
/D [27 0 R /FitR 30 699 297 672]
>>
endobj
52 0 obj
<<
/D [27 0 R /FitR 30 659 297 632]
>>
endobj
53 0 obj
<<
/D [27 0 R /FitR 30 639 297 612]
>>
endobj
54 0 obj
<<
/D [27 0 R /FitR 30 619 297 592]
>>
endobj
55 0 obj
<<
/D [27 0 R /FitR 30 589 297 562]
>>
endobj
56 0 obj
<<
/D [27 0 R /FitR 30 569 297 542]
>>
endobj
57 0 obj
<<
/D [27 0 R /FitR 30 549 297 522]
>>
endobj
58 0 obj
<<
/D [27 0 R /FitR 30 529 297 502]
>>
endobj
59 0 obj
<<
/D [26 0 R /FitR 298 477 565 450]
>>
endobj
60 0 obj
<<
/D [27 0 R /FitR 30 499 297 472]
>>
endobj
61 0 obj
<<
/D [27 0 R /FitR 30 479 297 452]
>>
endobj
62 0 obj
<<
/D [27 0 R /FitR 294 739 560 712]
>>
endobj
63 0 obj
<<
/D [27 0 R /FitR 294 709 560 682]
>>
endobj
64 0 obj
<<
/D [27 0 R /FitR 294 689 560 662]
>>
endobj
65 0 obj
<<
/D [27 0 R /FitR 294 649 560 622]
>>
endobj
66 0 obj
<<
/D [27 0 R /FitR 294 619 560 592]
>>
endobj
67 0 obj
<<
/D [27 0 R /FitR 294 599 560 572]
>>
endobj
68 0 obj
<<
/D [27 0 R /FitR 294 569 560 542]
>>
endobj
69 0 obj
<<
/D [27 0 R /FitR 294 559 560 532]
>>
endobj
70 0 obj
<<
/D [26 0 R /FitR 298 457 565 430]
>>
endobj
71 0 obj
<<
/D [27 0 R /FitR 294 539 560 512]
>>
endobj
72 0 obj
<<
/D [27 0 R /FitR 294 519 560 492]
>>
endobj
73 0 obj
<<
/D [27 0 R /FitR 294 489 560 462]
>>
endobj
74 0 obj
<<
/D [27 0 R /FitR 294 469 560 442]
>>
endobj
75 0 obj
<<
/D [26 0 R /FitR 298 437 565 410]
>>
endobj
76 0 obj
<<
/D [26 0 R /FitR 298 417 565 390]
>>
endobj
77 0 obj
<<
/D [26 0 R /FitR 298 387 565 360]
>>
endobj
78 0 obj
<<
/D [26 0 R /FitR 298 357 565 330]
>>
endobj
79 0 obj
<<
/D [26 0 R /FitR 298 337 565 310]
>>
endobj
80 0 obj
<<
/D [17 0 R /FitR 30 409 297 381]
>>
endobj
81 0 obj
<<
/D [18 0 R /FitR 35 497 301 469]
>>
endobj
82 0 obj
<<
/D [18 0 R /FitR 35 480 301 452]
>>
endobj
83 0 obj
<<
/D [18 0 R /FitR 35 310 301 282]
>>
endobj
84 0 obj
<<
/D [18 0 R /FitR 35 95 301 67]
>>
endobj
85 0 obj
<<
/D [18 0 R /FitR 298 453 565 425]
>>
endobj
86 0 obj
<<
/D [18 0 R /FitR 298 436 565 408]
>>
endobj
87 0 obj
<<
/D [19 0 R /FitR 294 625 560 597]
>>
endobj
88 0 obj
<<
/D [20 0 R /FitR 35 106 301 78]
>>
endobj
89 0 obj
<<
/D [21 0 R /FitR 294 739 560 711]
>>
endobj
90 0 obj
<<
/D [25 0 R /FitR 30 106 297 78]
>>
endobj
91 0 obj
<<
/D [19 0 R /FitR 30 541 560 40]
>>
endobj
92 0 obj
<<
/D [21 0 R /FitR 30 511 560 40]
>>
endobj
93 0 obj
<<
/D [22 0 R /FitR 35 734 565 247]
>>
endobj
94 0 obj
<<
/D [23 0 R /FitR 30 734 560 180]
>>
endobj
95 0 obj
<<
/D [24 0 R /FitR 35 734 565 180]
>>
endobj
96 0 obj
<<
/D [17 0 R /FitR 0 792 594 0]
>>
endobj
97 0 obj
<<
/D [20 0 R /FitR 35 100 301 87]
>>
endobj
98 0 obj
<<
/D [21 0 R /FitR 294 733 560 719]
>>
endobj
99 0 obj
<<
/D [25 0 R /FitR 30 100 297 87]
>>
endobj
100 0 obj
<<
/D [26 0 R /FitR 298 593 565 580]
>>
endobj
101 0 obj
<<
/D [26 0 R /FitR 298 547 565 534]
>>
endobj
102 0 obj
<<
/D [17 0 R /FitR 30 404 297 390]
>>
endobj
103 0 obj
<<
/D [18 0 R /FitR 35 491 301 477]
>>
endobj
104 0 obj
<<
/D [18 0 R /FitR 35 474 301 461]
>>
endobj
105 0 obj
<<
/D [18 0 R /FitR 35 304 301 290]
>>
endobj
106 0 obj
<<
/D [18 0 R /FitR 35 89 301 76]
>>
endobj
107 0 obj
<<
/D [18 0 R /FitR 298 447 565 433]
>>
endobj
108 0 obj
<<
/D [18 0 R /FitR 298 430 565 417]
>>
endobj
109 0 obj
<<
/D [19 0 R /FitR 294 619 560 606]
>>
endobj
110 0 obj
<<
/First 269 0 R
/Parent 9 0 R
/Next 270 0 R
/Dest
/Count -3
/Last 271 0 R
/Prev 15 0 R
/Title
>>
endobj
111 0 obj
<<
/Parent 9 0 R
/Next 16 0 R
/Dest
/Prev 272 0 R
/Title
>>
endobj
112 0 obj
<<
/Rect [111.345 610.469 120.869 620.05]
/Subtype /Link
/A <<
/URI (https://orcid.org/0000-0001-5210-650X)
/S /URI
>>
/Border [0 0 0]
/Type /Annot
>>
endobj
113 0 obj
<<
/Rect [358.129 610.469 367.71 620.05]
/Subtype /Link
/A <<
/URI (https://orcid.org/0000-0003-3733-3725)
/S /URI
>>
/Border [0 0 0]
/Type /Annot
>>
endobj
114 0 obj
<<
/Rect [202.053 359.376 220.309 368.901]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
115 0 obj
<<
/Rect [155.962 337.379 174.161 346.847]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
116 0 obj
<<
/Rect [218.211 337.379 236.466 346.847]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
117 0 obj
<<
/Rect [72.794 172.346 90.992 181.871]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
118 0 obj
<<
/Rect [488.579 364.876 506.211 374.4]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
119 0 obj
<<
/Rect [327.175 353.877 344.863 363.345]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
120 0 obj
<<
/Rect [531.723 320.882 549.411 330.35]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
121 0 obj
<<
/Rect [351.836 309.883 369.524 319.351]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
122 0 obj
<<
/Rect [327.005 298.885 344.693 308.353]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
123 0 obj
<<
/Rect [416.239 298.885 433.927 308.353]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
124 0 obj
<<
/Rect [494.475 298.885 512.107 308.353]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
125 0 obj
<<
/Rect [325.644 287.887 343.332 297.354]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
126 0 obj
<<
/Rect [513.241 276.888 530.929 286.356]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
127 0 obj
<<
/Rect [335.452 221.896 353.14 231.364]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
128 0 obj
<<
/Rect [435.855 221.896 453.486 231.364]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
129 0 obj
<<
/Rect [518.23 221.896 535.918 231.364]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
130 0 obj
<<
/Rect [324.113 210.898 341.801 220.365]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
131 0 obj
<<
/Rect [407.112 210.898 424.743 220.365]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
132 0 obj
<<
/Rect [370.375 199.899 388.063 209.367]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
133 0 obj
<<
/Rect [338.06 177.846 355.748 187.37]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
134 0 obj
<<
/Rect [423.496 177.846 441.184 187.37]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
135 0 obj
<<
/Rect [531.723 166.847 549.354 176.372]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
136 0 obj
<<
/Rect [377.235 155.849 394.923 165.373]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
137 0 obj
<<
/Rect [485.745 133.852 503.433 143.376]
/Subtype /Link
/Border [0 0 0]
/Dest
/Type /Annot
>>
endobj
138 0 obj
<<
/Rect [200.183 125.688 287.887 132.718]
/Subtype /Link
/A <<
/URI (mailto:jwillemse@biology.leidenuniv.nl)
/S /URI
>>
/Border [0 0 0]
/Type /Annot
>>
endobj
139 0 obj
<<
/Rect [48.926 89.688 129.94 96.718]
/Subtype /Link
/A <<
/URI (https://doi.org/10.1017/S1431927620024320)
/S /URI
>>
/Border [0 0 0]
/Type /Annot
>>
endobj
140 0 obj
<<
/Rect [208.913 63.78 348.775 70.809]
/Subtype /Link
/A <<
/URI (http://creativecommons.org/licenses/by-nc-sa/4.0/)
/S /URI
>>
/Border [0 0 0]
/Type /Annot
>>
endobj
141 0 obj
<<
/Type /Annot
/Subtype /Link
/Rect [524.57477 8.0 583.972 30.845076]
/Border [0 0 0]
/A 273 0 R
>>
endobj
142 0 obj
<<
/Type /Annot
/Subtype /Link
/Rect [7.7999997 14.264648 133.62 22.189453]
/Border [0 0 0]
/A 274 0 R
>>
endobj
143 0 obj
<<
/Filter /FlateDecode
/Length 10
>>
stream
x+ |
endstream
endobj
144 0 obj
<<
/Filter /FlateDecode
/Length 10
>>
stream
x+ |
endstream
endobj
145 0 obj
<<
/Length 2155
/Filter /FlateDecode
>>
stream
HWێF}Wpݼ>q# 6A`E^SLR3V~Tb=E.N]SUN"#"ߔeA_5|_