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This chapter examines the concept of communication traditionally and currently used in the field of ethology. It describes a few landmarks in the evolution of communication in the different species, paying particular on the mechanisms that regulate these features since this is where the levels of progressive complexity of processing are found. The different modalities of communication have evolved to serve the general function of regulating the (social) behavior of each species within its own ecological niche. The emergence of birds and mammals was accompanied by a reorganization of perceptive systems, which were for the first-time centralized in the brain. The chapter focuses on communication in primates, and in particular the anthropoids, since these animals make use of forms of communication very close to those of humans. Finally, the chapter focuses on describes the sign created by the hominid mind in order to facilitate communication between minds.