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 This research was supported by funds from Emory University, Children's Healthcare of Atlanta, the Emory Center for AIDS Research, the Georgia Research Alliance, Human Frontiers Science Program, National Institutes of Health (1R01GM104540 and 1R21AI101775), and the National Science Foundation (0923395) to E.R.W, and S10 RR025679 to P.W.S. Cryo-EM/ET data was collected at the Emory University Robert P. Apkarian Integrated Electron Microscopy Core..