Hostname: page-component-8448b6f56d-c47g7 Total loading time: 0 Render date: 2024-04-24T13:50:21.156Z Has data issue: false hasContentIssue false

A restudy of the Sandbian to Katian (Upper Ordovician) graptolites from the East Qilianshan (Chilianshan), Northwest China

Published online by Cambridge University Press:  22 August 2019

Xu Chen*
Affiliation:
CAS Key Laboratory of Economic Stratigraphy and Palaeogeography, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China Academician Studio, Zhejiang Nuclear Industry Geological Survey Team, Huzhou 313000, China
Zhongyang Chen
Affiliation:
Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China
Charles E. Mitchell
Affiliation:
Department of Geology, University at Buffalo, State University of New York, 126 Cooke Hall, Buffalo, New York 14260, USA
Qing Chen
Affiliation:
CAS Key Laboratory of Economic Stratigraphy and Palaeogeography, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China
Linna Zhang
Affiliation:
State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China
*
*Corresponding author

Abstract

The Upper Ordovician Sandbian to Katian strata from the East Qilianshan (northeastern Tibet Plateau) bear a graptolite fauna of moderately high diversity. Graptolites from the Amplexograptus maxwelli beds to the Appendispinograptus longispinus Biozone (Sa2–Ka4 intervals) proposed herein include 27 species of 13 genera. This important graptolite fauna is first described herein although it was initially reported in 1963. Most of them occur in the A. longispinus Biozone corresponding to the Dicellograptus complexus to Paraorthograptus pacificus biozones of the Wufeng Formation in the Yangtze region. Alulagraptus new genus is established based on the materials from the East Qiqiaogou section. The endemic species, e.g., Alulagraptus ensiformis (Mu and Zhang in Mu et al., 1963), Dicellograptus sinicus Mu and Zhang in Mu et al., 1963, and Climacograptus? papilio Mu and Zhang in Mu et al., 1963, could indicate that East Qilianshan block was separated from South China.

UUID: http://zoobank.org/84ab69a2-0b56-4ec3-912e-730da08ab10a

Type
Articles
Copyright
Copyright © 2019, The Paleontological Society 

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Bates, D.E.B., and Kirk, N.H., 1991, The ultrastructure, mode of construction and functioning of Ordovician retiolitid graptolites from the Viola Springs Limestone, Oklahoma: Modern Geology, v. 15, p. 131286.Google Scholar
Bergström, S.M., and Mitchell, C.E., 1990, Trans-Pacific graptolite-faunal relations: The biostratigraphic position of the base of the Cincinnatian Series (Upper Ordovician) in the standard Australian graptolite zone succession: Journal of Paleontology, v. 64, p. 992997.Google Scholar
Bergström, S.M., Chen, X., Gutiérrez-Marco, J.C., and Dronov, A., 2009, The new chronostratigraphic classification of the Ordovician system and its relations to major regional series and stages and to δ13C chemostratigraphy: Lethaia, v. 42, p. 97107, doi:10.1111/j.1502-3931.2008.00136.x.Google Scholar
Berry, W.B.N., 1960, Graptolite faunas of the Marathon region, west Texas: University of Texas Publication, v. 6005, p. 1179.Google Scholar
Berry, W.B.N., 1966, A discussion of some Victorian Ordovician graptolites: Proceedings of the Royal Society of Victoria, v. 79, p. 415448.Google Scholar
Bolton, T.E., 1960, Catalogue of Type Invertebrate Fossils of the Geological Survey of Canada, Volume 1: Ottawa, Geological Survey of Canada, 215 p.Google Scholar
Boucot, A.J., Chen, X., and Scotese, C.R., 2013, Phanerozoic paleoclimate: An atlas of lithologic indicators of climate: SEPM Concepts in Sedimentology and Paleontology, v. 11, p. 1478, doi:10.2110/sepmcsp.11.Google Scholar
Boyle, J., Sheets, H.D., Wu, S.Y., Goldman, D., Melchin, M.J., Cooper, R.A., Sadler, P.M., and Mitchell, C.E., 2017, The impact of geographic range, sampling, ecology, and time on extinction risk in the volatile clade Graptoloida: Paleobiology, v. 43, p. 85113, doi:10.1017/pab.2016.33.Google Scholar
Bulman, O.M.B., 1931, South American graptolites with special reference to the Nordenskiold Collection: Arkiv för Zoologi, v. 22A, p. 1111, pls. 1–12.Google Scholar
Bulman, O.M.B., 1945–1947, A monograph of the Caradoc (Balclatchie) graptolites from limestones in Laggan Burn, Ayrshire, Parts 1–3: Palaeontographical Society, v. 98, p. 142, pls. 1–3 (1945); p. 43–58, pls. 4–6 (1946); p. 59–78, pls. 7–10 (1947).Google Scholar
Carruthers, W., 1867, Graptolites: Their structure and systematic position: Intellectual Observer, v. 11, p. 283292, 365–374.Google Scholar
Carter, C., 1972, Ordovician (upper Caradocian) graptolites from Idaho and Nevada: Journal of Paleontology, v. 46, p. 4349.Google Scholar
Carter, C., and Churkin, M., 1977, Ordovician and Silurian graptolite succession in the Trail Creek area, central Idaho—A graptolite zone reference section: United States Geological Survey Professional Paper, v. 1020, p. 137.Google Scholar
Chen, X., Ni, Y.N., Mitchell, C.E., Qiao, X.D., and Zhan, S.G., 2000a, Graptolites from the Qilang and Yingan formations (Caradoc, Ordovician) of Kalpin, western Tarim, Xinjiang, China: Journal of Paleontology, v. 74, p. 282300, doi:10.1666/0022-3360(2000)074<0282:GFTQAY > 2.0.CO;2.+2.0.CO;2.>Google Scholar
Chen, X., Rong, J.Y., Mitchell, C.E., Harper, D.A.T., Fan, J.X., Zhan, R.B., Zhang, Y.D., Li, R.Y., and Wang, Y., 2000b, Late Ordovician to earliest Silurian graptolite and brachiopod biozonation from the Yangtze region, South China with a global correlation: Geological Magazine, v. 137, p. 623650, doi:10.1017/S0016756800004702.Google Scholar
Chen, X., Fan, J.X., Melchin, M.J., and Mitchell, C.E., 2005, Hirnantian (latest Ordovician) graptolites from the Yangtze region, China: Palaeontology, v. 48, p. 235280, doi:10.1111/j.147504983.2005.00453.x.Google Scholar
Chen, X., Zhang, Y.D., Li, Y., Fan, J.X., Tang, P., Chen, Q., and Zhang, Y.Y., 2012, Biostratigraphic correlation of the Ordovician black shales in Tarim Basin and its peripheral regions: Science China Earth Sciences, v. 55, p. 12301237, doi:10.1007/s11430-012-4448-6.Google Scholar
Chen, X., Zhang, Y.D., Goldman, D., Bergström, S.M., Fan, J.X., Wang, Z.H., Finney, S.C., Chen, Q., and Ma, X., 2017, Darriwilian to Katian (Ordovician) Graptolites from Northwest China: Amsterdam, Elsevier, 370 p.Google Scholar
Cocks, L.R.M., and Torsvik, T.H., 2013, The dynamic evolution of the Palaeozoic geography of eastern Asia: Earth-Science Reviews, v. 117, p. 4079, doi:10.1016/j.earscirev.2012.12.001.Google Scholar
Cooper, R.A., and Sadler, P., 2010, Facies preference predicts extinction risk in Ordovician graptolites: Paleobiology, v. 32, p. 167187, doi:10.1666/09043.1.Google Scholar
Davies, K.A., 1929, Notes on the graptolite faunas of the Upper Ordovician and lower Silurian: Geological Magazine, v. 66, p. 127.Google Scholar
Decker, C.E., 1935, The graptolites of the Simpson Group of Oklahoma: Proceedings of the National Academy of Sciences of the United States of America, v. 21, p. 239243.Google Scholar
Elles, G.L., and Wood, E.M.R., 1904, A monograph of British graptolites, Part 4: Monographs of the Palaeontographical Society, v. 58, p. liiilxii, 135–180.Google Scholar
Elles, G.L., and Wood, E.M.R., 1906, A monograph of British graptolites, Part 5: Monographs of the Palaeontographical Society, v. 60, p. lxxiiixcvi, 181–216.Google Scholar
Elles, G.L., and Wood, E.M.R., 1907, A monograph of British graptolites, Part 6: Monographs of the Palaeontographical Society, v. 61, p. xcviicxx, 217–272.Google Scholar
Elles, G.L., and Wood, E.M.R., 1908, A monograph of British graptolites, Part 7: Monographs of the Palaeontographical Society, v. 62, p. cxxicxlviii, 273–358.Google Scholar
Finney, S.C., 1986, Graptolite biofacies and correlation of eustatic, subsidence, and tectonic events in the Middle to Upper Ordovician of North America: PALAIOS, v. 1, p. 435461.Google Scholar
Frech, F., 1897, Lethaea Geognostica 1, Lethaea Palaeozoica 1, Graptolithen: Stuttgart, Schweizerbart, 690 p.Google Scholar
Fu, H.Y., 1982, Graptolithina, in Geological Bureau of Hunan, ed., The Palaeontological Atlas of Hunan: Beijing, Geological Publishing House, p. 410479. [in Chinese with English abstract]Google Scholar
Ge, M.Y., 1984, [Upper Ordovician graptolites from Kunshan, Jiangsu]: Bulletin of Nanjing Institute of Geology and Palaeontology, Academia Sinica, v. 3, p. 237263. [in Chinese with English abstract]Google Scholar
Goldman, D., 1995, Taxonomy, evolution, and biostratigraphy of the Orthograptus quadrimucronatus species group (Ordovician, Graptolithina): Journal of Paleontology, v. 69, p. 516540.10.1017/S0022336000034909Google Scholar
Goldman, D., and Bergström, S.M., 1997, Late Ordovician graptolites from the North American midcontinent: Palaeontology, v. 40, p. 9651010.Google Scholar
Goldman, D., Bergström, S.M., and Mitchell, C.E., 1995, Revision of the Zone 13 graptolite biostratigraphy in the Marathon, Texas, standard succession and its bearing on Upper Ordovician graptolite biogeography: Lethaia, v. 28, p. 115128.Google Scholar
Goldman, D., Campbell, S.M., and Rahl, J.M., 2002, Three-dimensionally preserved specimens of Amplexograptus (Ordovician, Graptolithina) from the North American mid-continent: Taxonomic and biostratigraphic significance: Journal of Paleontology, v. 76, p. 921927, doi:10.1666/0022-3360(2002)076<0921:TDPSOA>2.0.CO;2.2.0.CO;2.>Google Scholar
Goldman, D., Leslie, S.A., Nõlvak, J., Young, S., and Bergström, S.M., 2007, The Global Stratotype Section and Point (GSSP) for the base of the Katian Stage of the Upper Ordovician Series at Black Knob Ridge, southeastern Oklahoma, USA: Episodes, v. 30, p. 258270.Google Scholar
Goldman, D., Mitchell, C.E., Melchin, M.J., Fan, J.X., Wu, S.Y., and Sheets, H.D., 2011, Biogeography and mass extinction: Extirpation and re-invasion of Normalograptus species (Graptolithina) in the Late Ordovician palaeotropics: Proceedings of the Yorkshire Geological Society, v. 58, p. 227246, doi:10.1144/Pygs.58.4.300.Google Scholar
Gradstein, F., Ogg, J., Schmitz, M., and Ogg, G., 2012, The Geological Time Scale 2012: Oxford, UK, Elsevier, 1144 p.Google Scholar
Hall, J., 1847, Palaeontology of New York, Volume 1: Descriptions of the Organic Remains of the Lower Division of the New-York System: Albany, New York, C. Van Benthuysen, 338 p.Google Scholar
Hall, J., 1859, Notes on the genus Graptolithus, in Hall, J., ed., Palaeontology, Volume 3, Containing Descriptions and Figures of the Organic Remains of the Lower Helderberg Group and the Oriskany Sandstone, 1855–1859: Albany, New York, C. Van Benthuysen, p. 495522.Google Scholar
Hall, J., 1865, Graptolites of the Quebec Group, figures and descriptions of Canadian organic remains: Canadian Geological Survey Decade, v. 2, p. 1151.Google Scholar
Hall, T.S., 1902, The graptolites of New South Wales, in the collection of the Geological Survey: Records of the Geological Survey of New South Wales, v. 7, p. 4959.Google Scholar
Herr, S.R., 1971, Regeneration and growth abnormalities in Orthograptus quadrimucronatus from the Ordovician Maquoketa Formation of Iowa: Journal of Paleontology, v. 45, p. 628632.Google Scholar
Hopkinson, J., 1871, On Dicellograptus, a new genus of graptolites: Geological Magazine, v. 8, p. 2026.Google Scholar
Hsü, S.C., 1959, [A new graptolite fauna from the Lower Ordovician shale of Tsaidam, Chinghai Province]: Acta Palaeontologica Sinica, v. 7, p. 161192. [in Chinese with English summary]Google Scholar
Jacobi, R.D., and Mitchell, C.E., 2018, Aseismic-ridge subduction as a driver for the Ordovician Taconic orogeny and Utica foreland basin in New England and New York State: Geological Society of America Special Papers, v. 540, p. 617659, doi:10.1130/2018.2540(27).Google Scholar
Keble, R.A., and Harris, W.J., 1934, Graptolites of Victoria: New species and additional records: Memoirs of the National Museum, Melbourne, v. 8, p. 166183.Google Scholar
Keller, B.M., 1956, [Graptolity Ordovika Chu-Iliyskikh Gor]: Akademiya Nauk SSSR, Trudy Geologicheskogo Instituta, v. 1, p. 50102. [in Russian]Google Scholar
Koren’, T.N., 1979, [Pacificograptus, a new Late Ordovician diplograptid genus]: Paleontological Journal, v. 1979, p. 6570. [in Russian]Google Scholar
Koren’, T.N., Sobolevskaya, R.F., Mikhaylova, N.F., and Tsai, D.T., 1979, New evidence on graptolite succession across the Ordovician-Silurian Boundary in the Asian Part of the USSR: Acta Palaeontologica Polonica, v. 24, p. 125136.Google Scholar
Koren’, T.N., Mikhaylova, N.F., and Tsay, D.T., 1980, [Klass Graptolithina, Graptolity], in Apollonov, M.K., Bandaletov, S.M., and Nikitin, I.F., eds., [The Ordovician–Silurian Boundary in Kazakhstan]: Alma-Ata, Kazakhstan, Kazakh SSR Publishing House, p. 121170. [in Russian]Google Scholar
Koren’, T.N., Oradovskaya, M.M., Pylma, L.J., Sobolevskaya, R.F., and Chugaeva, M.N., 1983, [The Ordovician and Silurian Boundary in the Northeast of USSR]: Leningrad, Nauka Publishers, 192 p. [in Russian with English abstract]Google Scholar
Kraft, P., Štorch, P., and Mitchell, C.E., 2015, Graptolites of the Králův Dvůr Formation (mid Katian to earliest Hirnantian, Czech Republic): Bulletin of Geosciences, v. 90, p. 195225, doi:10.3140/bull.geosci.1435.Google Scholar
Lapworth, C., 1873, Notes on the British graptolites and their allies: Geological Magazine, v. 10, p. 500504, 555–560.Google Scholar
Lapworth, C., 1876, On Scottish Monograptidae: Geological Magazine, v. 13, p. 308321, 350–360, 499–507, 544–552.Google Scholar
Lapworth, C., 1877, On the graptolites of County Down: Proceedings of the Belfast Naturalists’ Field Club, v. 1876–1877, p. 125148.Google Scholar
Lapworth, C., 1880, On new British graptolites: Annals and Magazine of Natural History, v. 5, p. 149177.Google Scholar
Li, J.J., 1984, [Graptolites from the Xinling Formation (Upper Ordovician) of South Anhui]: Memoirs of Nanjing Institute of Geology and Palaeontology, Academia Sinica, v. 20, p. 145194. [in Chinese with English abstract]Google Scholar
Li, Y.F., Schieber, J., Fan, T.L., Li, Z.Y., and Zhang, J.P., 2017, Regional depositional changes and their controls on carbon and sulfur cycling across the Ordovician-Silurian boundary, northwestern Guizhou, South China: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 485, p. 816832, doi:10.1016/j.palaeo.2017.07.039.Google Scholar
Li, Z.M., and Li, D.Q., 1985, [Appendispinograptus, a new subgenus of Climacograptus]: Earth Science Journal of Wuhan College of Geology, v. 10, p. 3542. [in Chinese with English abstract]Google Scholar
Loxton, J., Melchin, M.J., Mitchell, C.E., and Senior, S.J.H., 2011, Ontogeny and astogeny of the graptolite genus Appendispinograptus (Li and Li, 1985): Proceedings of the Yorkshire Geological Society, v. 58, p. 253260, doi:10.1144/pygs.58.4.309.Google Scholar
Loxton, J.D., 2017, Graptolite diversity and community changes surrounding the Late Ordovician Mass Extinction: High resolution data from the Blackstone River, Yukon [Ph.D. thesis]: Halifax, Nova Scotia, Dalhousie University, 590 p.Google Scholar
Loydell, D., 2012, Graptolite biozone correlation charts: Geological Magazine, v. 149, p. 124132, doi:10.1017/S0016756811000513.Google Scholar
Maletz, J., 2014, The classification of the Pterobranchia (Cephalodiscida and Graptolithina): Bulletin of Geosciences, v. 89, p. 477540, doi:10.3140/bull.geosci.1465.Google Scholar
Maletz, J., Carlucci, J., and Mitchell, C.E., 2009, Graptoloid cladistics, taxonomy and phylogeny: Bulletin of Geosciences, v. 84, p. 719, doi:10.3140/bull.geosci.1108.Google Scholar
Maletz, J., Bates, D.E.B., Brussa, E.D., Cooper, R.A., Lenz, A.C., Riva, J.F., Toro, B.A., and Zhang, Y.D., 2014, Glossary of the Hemichordata, in Treatise [on Invertebrate Paleontology] Online, Part V (Revision 2), Chapter 12, no. 62, 23 p. [see https://journals.ku.edu/treatiseonline/issue/view/398].Google Scholar
Melchin, M.J., 1987, Upper Ordovician graptolites from the Cape Phillips Formation, Canadian Artic Islands: Bulletin of the Geological Society of Denmark, v. 35, p. 191202.Google Scholar
Melchin, M.J., Holmden, C., and Williams, S.H., 2003, Correlation of graptolite biozones, chitinozoan biozones, and carbon isotope curves through the Hirnantian: INSUEGO, Serie Correlación Geológica, v. 17, p. 101104.Google Scholar
Mitchell, C.E., 1987, Evolution and phylogenetic classification of the Diplograptacea: Palaeontology, v. 30, p. 353405.Google Scholar
Mitchell, C.E., 1988, The morphology and ultrastructure of Brevigraptus quadrithecatus n. gen., n. sp. (Diplograptacea), and its convergence upon Dicaulograptus hystrix (Bulman): Journal of Paleontology, v. 62, p. 448463.Google Scholar
Mitchell, C.E., Goldman, D., Cone, M.R., Maletz, J., and Janousek, H., 2003, Ordovician graptolites of the Phi Kappa Formation at Trail Creek, central Idaho, USA: A revised biostratigraphy: INSUGEO, Serie Correlación Geológica, v. 18, p. 6972.Google Scholar
Mitchell, C.E., Chen, X., and Finney, S.C., 2007a, The structure and possible function of ‘basal membranes’ in the spinouse climacograptid graptolite Appendispinograptus Li and Li 1985: Journal of Paleontology, v. 81, p. 11221127, doi:10.1666/pleo05-105.1.Google Scholar
Mitchell, C.E., Goldman, D., Klosterman, S.L., Maletz, J., Sheets, H.D., and Melchin, M.J., 2007b, Phylogeny of the Ordovician Diplograptoidea: Acta Palaeontologica Sinica, v. 46, p. 332339.Google Scholar
Mu, E.Z., 1963a, [On the complication of graptolite rhabdosome]: Acta Palaeontologica Sinica, v. 11, p. 346377. [in Chinese with English summary]Google Scholar
Mu, E.Z., 1963b, Research in graptolite fauna of Chilianshan: Scientia Sinica, v. 12, p. 347371.Google Scholar
Mu, E.Z., 1974, Evolution, classification and distribution of Graptoloidea and Graptodendroids: Scientia Sinica, v. 17, p. 227238.Google Scholar
Mu, E.Z., and Lin, Y.K., 1984, Graptolites from the Ordovician-Silurian boundary sections of Yichang area, W. Hubei, in Nanjing Institute of Geology and Palaeontology, Academia Sinica, ed., Stratigraphy and Palaeontology of Systemic Boundaries in China, Ordovician-Silurian Boundary 1: Hefei, China, Anhui Science and Technology Publishing House, p. 4573.Google Scholar
Mu, E.Z., and Zhang, Y.K., 1964, [Ordovician and Silurian graptolite bearing strata from the eastern Qilianshan (Chilianshan)]: Memoirs of Nanjing Institute of Geology and Palaeontology, Academia Sinica, Stratigraphy Special Issue, v. 1, p. 120. [in Chinese]Google Scholar
Mu, E.Z., Li, J.J., Ge, M.Y., Chen, X., and Zhang, Y.K., 1963, [Ordovician, Late Ordovician, graptolites], in Institute of Geology and Palaeontology, Academia Sinica, ed., [A Handbook of Index Fossils of North-west China]: Beijing, Science Press, p. 5760. [in Chinese]Google Scholar
Mu, E.Z., Li, J.J., Ge, M.Y., Chen, X., Ni, Y.N., Lin, Y.K., and Mu, X.N., 1974, [Ordovician, Graptolites], in Nanjing Institute of Geology and Palaeontology, Academia Sinica, ed., [A Handbook of Stratigraphy and Palaeontology of South-west China]: Beijing, Science Press, p. 154164. [in Chinese]Google Scholar
Mu, E.Z., Song, L.S., Li, J.S., Xu, B.Z., and Zhang, Y.K., 1982, [Graptolithina], in Xi'an Institute of Geology and Mineral Resources, ed., [Palaeontological Atlas of Northwest China (Shaanxi, Gansu and Ningxia), 1, Precambrian to early Paleozoic]: Beijing, Geological Publishing House, p. 294346. [in Chinese]Google Scholar
Mu, E.Z., Li, J.J., Ge, M.Y., Chen, X., Lin, Y.K., and Ni, Y.N., 1993, [Upper Ordovician graptolites of central China region]: Palaeontologia Sinica, new ser. B, v. 29, p. 1393. [in Chinese with English summary]Google Scholar
Mu, E.Z., Li, J.J., Ge, M.Y., Lin, Y.K., and Ni, Y.N., 2002, [Fossil Graptolites of China]: Beijing, Science Press, 1205 p. [in Chinese]Google Scholar
Nicholson, H.A., 1869, On some new species of graptolites: Annals and Magazine of Natural History, v. 4, p. 231242.Google Scholar
Perner, J., 1895, Études sur les Graptolites de Bohême, IIième Partie: Monographie des Graptolites de l’Étage D: Prague, Raimund Gerhard, 31 p.Google Scholar
Přibyl, A., 1947, [On the classification of the genus Climacograptus Hall, 1865]: Bulletin Internationale de l'Académie Tchèque des Sciences, v. 48, p. 112.Google Scholar
Přibyl, A., 1949, [Revision of the Diplograptidae and Glossograptidae of the Ordovician of Bohemia]: Bulletin Internationale de l'Académie Tchèque des Sciences, v. 50, p. 151.Google Scholar
Riva, J., 1974a, A revision of some Ordovician graptolites of eastern North America: Palaeontology, v. 17, p. 140.Google Scholar
Riva, J., 1974b, Late Ordovician spinose climacograptids from the Pacific and Atlantic faunal provinces: Special Papers in Palaeontology, v. 13, p. 107126.Google Scholar
Riva, J., 1987, The graptolite Amplexograptus praetypicalis n. sp. and the origin of the typicalis group: Canadian Journal of Earth Sciences, v. 24, p. 924933.Google Scholar
Riva, J., 1988, Graptolites at and below the Ordovician-Silurian boundary on Anticosti Island, Canada: Bulletin of the British Museum (Natural History), Geology, v. 43, p. 221237.Google Scholar
Ross, R.J.J., and Berry, W.B.N., 1963, Ordovician graptolites of the Basin Ranges in California, Nevada, Utah, and Idaho: United States Geological Survey Bulletin, v. 1134, p. 1177.Google Scholar
Ruedemann, R., 1908, Graptolites of New York, Part II: Memoirs of New York State Museum, v. 11, p. 1481.Google Scholar
Ruedemann, R., 1912, The Lower Siluric shales of the Mohawk Valley: Bulletin of New York State Museum, v. 162, p. 1145.Google Scholar
Ruedemann, R., 1947, Graptolites of North America: Geological Society of America Memoir, v. 19, p. 1652.Google Scholar
Sheets, H.D., Mitchell, C.E., Melchin, M.J., Loxton, J., Štorch, P., Carlucci, K.L., and Hawkins, A.D., 2016, Graptolite community responses to global climate change and the Late Ordovician mass extinction: Proceedings of the National Academy of Sciences, v. 113, p. 83808385, doi:10.1073/pnas.1602102113.Google Scholar
Song, S.G., Niu, Y.L., Su, L., and Xia, X.H., 2013, Tectonics of the North Qilian orogen, NW China: Gondwana Research, v. 23, p. 13781401, doi:10.1016/j.gr.2012.02.004.Google Scholar
Stewart, S., and Mitchell, C.E., 1997, Anticostia, a distinctive new Late Ordovician ‘glyptograptid’ (Diplograptacea, Graptoloidea) based on three-dimensionally preserved specimens from Anticosti Island, Quebec: Canadian Journal of Earth Sciences, v. 34, p. 215228.Google Scholar
Štorch, P., Mitchell, C.E., Finney, S.C., and Melchin, M.J., 2011, Uppermost Ordovician (upper Katian–Hirnantian) graptolites of north-central Nevada, U.S.A.: Bulletin of Geosciences, v. 86, p. 301386, doi:10.3140/bull.geosci.1264.Google Scholar
Thomas, D.E., 1960, The zonal distribution of Australian graptolites: Journal and Proceedings of the Royal Society of New South Wales, v. 94, p. 158.Google Scholar
Toghill, P., 1970, A fauna from the Hendre Shales (Llandeilo) of the Mydrim area, Carmarthenshire: Proceedings of the Geological Society of London, v. 1663, p. 121129.Google Scholar
Torsvik, T.H., and Cocks, L.R.M., 2017, Earth History and Palaeogeography: Cambridge, UK, Cambridge University Press, 317 p.Google Scholar
VandenBerg, A.H.M., 2018, Climcaograptus? uncinatus Keble & Harris, 1934, in Wilkinson, J.C., Zalasiewicz, J.A., Rushton, A.W.A., and Muir, L.A., eds., Atlas of Graptolite Type Specimens, Folio 3: Palaeontographical Society and British and Irish Graptolite Group, http://graptolites.com/atlas-folio/4590515350 (accessed July 2019), p. 3.93.Google Scholar
VandenBerg, A.H.M., and Cooper, R.A., 1992, The Ordovician graptolite sequence of Australasia: Alcheringa, v. 16, p. 3385.Google Scholar
Wang, X.F., Zeng, Q.L., Zhou, T.M., Ni, S.Z., Xu, G.H., Sun, Q.Y., Li, Z.H., Xiang, L.W., and Lai, C.G., 1983, [Latest Ordovician and earliest Silurian faunas from the eastern Yangtze gorges, China, with comments on Ordovician-Silurian boundary]: Bulletin of the Yichang Institute of Geology and Mineral Resources, Academy of Geological Sciences, v. 6, p. 95163. [in Chinese with English abstract]Google Scholar
Whittington, H.B., 1954, A new Ordovician graptolite from Oklahoma: Journal of Paleontology, v. 28, p. 613621.Google Scholar
Whittington, H.B., and Rickards, R.B., 1969, Development of Glossograptus and Skiagraptus, Ordovician graptoloids from Newfoundland: Journal of Paleontology, v. 43, p. 800817.Google Scholar
Williams, S.H., 1982, The late Ordovician graptolite fauna of the Anceps bands at Dob`s Linn, southern Scotland: Geologica et Palaeontologica, v. 16, p. 2956.Google Scholar
Williams, S.H., 1987, Upper Ordovician graptolites from the D. complanatus Zone of the Moffat and Girvan districts and their significance for correlation: Scottish Journal of Geology, v. 23, p. 6592.Google Scholar
Xia, G.S., 1982, [Graptolites of Anhui Province]: Hefei, China, Anhui Science and Technology Publishing House, 166 p. [in Chinese]Google Scholar
Yang, D.Q., Ni, Y.N., Li, J.J., Chen, X., Lin, Y.K., Yu, J.H., Xia, G.S., Jiao, S.D., Fang, Y.T., Ge, M.Y., and Mu, E.Z., 1983, [Graptolithina], in Nanjing Institute of Geology and Mineral Resources, ed., [Palaeontological Atlas of East China, 1, Early Paleozoic]: Beijing, Geological Publishing House, p. 353508. [in Chinese]Google Scholar
Yang, L.M., Song, S.G., Allen, M.B., Su, L., Dong, J.L., and Wang, C., 2018, Oceanic accretionary belt in the West Qinling Orogen: Links between the Qinling and Qilian orogens, China: Gondwana Research, v. 64, p. 137162, doi:10.1016/j.gr.2018.06.009.Google Scholar
Zalasiewicz, J.A., Rushton, A.W.A., and Owen, A.W., 1995, Late Caradoc graptolitic faunal gradients across the Iapetus Ocean: Geological Magazine, v. 132, p. 611617.Google Scholar
Zalasiewicz, J.A., Taylor, L., Rushton, A.W.A., Loydell, D.K., Rickards, R.B., and Williams, M., 2009, Graptolites in British stratigraphy: Geological Magazine, v. 146, p. 785850, doi:10.1017/S0016756809990434.Google Scholar
Zhao, J.H., Jin, Z.J., Jin, Z.K., Geng, Y.K., Wen, X., and Yan, C.N., 2016, Applying sedimentary geochemical proxies for paleoenvironment interpretation of organic-rich shale deposition in the Sichuan Basin, China: International Journal of Coal Geology, v. 163, p. 5271, doi:10.1016/j.coal.2016.06.015.Google Scholar