Hostname: page-component-7c8c6479df-ws8qp Total loading time: 0 Render date: 2024-03-19T04:38:13.065Z Has data issue: false hasContentIssue false

Lowermost Silurian graptolites from the Kurama Range, eastern Uzbekistan

Published online by Cambridge University Press:  20 May 2016

Tatjana N. Koren'
Affiliation:
All-Russia Geological Research Institute (VSEGEI), Sredny Pr. 74, 199026, St. Petersburg, Russia
Michael J. Melchin
Affiliation:
Geology Department, St. Francis Xavier University, P.O. Box 5000, Antigonish, Nova Scotia B2G 2W5, Canada

Abstract

An unusually diverse fauna of biserial graptolites has been recovered from the Kajrak and Urubulak formations in the Kurama Range of eastern Uzbekistan. The fauna consists of 26 distinct taxa, 24 of which are systematically described and illustrated. The entire fauna can be assigned to the Akidograptus ascensus–Parakidograptus acuminatus Biozone, which is earliest Rhuddanian (early Llandovery) in age. Within this biozone, however, three regionally distinguishable subzones can be recognized, the N. lubricus, A. cuneatus and H. sinitzini subzones. New taxa described are Normalograptus larini, Normalograptus skeliphrus, “Paraclimacograptus” kiliani kuramaensis, “Glyptograptus” jerini, and “Glyptograptus” ugurensis. New combinations described are Normalograptus acceptus (Koren’ and Mikhaylova, 1980), Normalograptus bicaudatus (Chen and Lin, 1978), Normalograptus jideliensis (Koren’ and Mikhaylova, 1980), Normalograptus lubricus (Chen and Lin, 1978), Normalograptus mirnyensis (Obut and Sobolevskaya, 1967), and Sudburigraptus illustris (Koren’ and Mikhaylova, 1980).

Type
Research Article
Copyright
Copyright © The Paleontological Society

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Biske, J. S. 1991. Volcanic arches in the Paleozoic history of the South Tien Shan. Geotektonika, 2:4146. (In Russian)Google Scholar
Bjerreskov, M. 1975. Llandoverian and Wenlockian graptolites from Bornholm. Fossils and Strata, 8:194.Google Scholar
Bulman, O. M. B. 1933. Programme evolution in the graptolites. Biological Reviews, 3:311333.CrossRefGoogle Scholar
Bulman, O. M. B. 1955. Graptolithina, p.1101. In Moore, R. C. (ed.). Treatise on Invertebrate Paleontology (V). Geological Society of America and University of Kansas Press, Lawrence.Google Scholar
Bulman, O. M. B., and Rickards, R. B. 1968. Some new diplograptids from the Llandovery of Britain and Scandinavia. Palaeontology, 11:115.Google Scholar
Burtman, V. S. 1976. Structural Evolution of Paleozoic Fold Systems. Nauka, Moscow, 164 p. (In Russian)Google Scholar
Carruthers, W. A. 1868. Revision of the British graptolites, with descriptions of the new species and notes on their affinities. Geological Magazine, 5:6474, 125–133.CrossRefGoogle Scholar
Chaletskaya, O. N. 1960. New species of the Llandovery graptolites of Central Asia, p. 373375. In Markovsky, B. P. (ed.), Novye Vidy Drevnikh Rastenii I Bespozvononochnykh SSSR, Chast’ II Moskva, Gosudarstvennoe Nauchno-Teknicheskoje Izdatel'stvo. (In Russian)Google Scholar
Chaletskaya, O. N. 1962. Llandovery graptolites of western Tien Shan, p. 54105. In Verkhov, V. J. (ed.), Stratigraphia and Paleontologia Uzbekistana I Sopredel'nykh Regionov, Kn. 1. Institut Geologii Akademii Nauk UzSSR. Tashkent. (In Russian)Google Scholar
Chen, Xu, and Yaokun, Lin, 1978. Lower Silurian graptolites from Tongzi, northern Guizhou. Memoirs of Nanjing Institute of Geology and Palaeontology, Academia Sinica, Number 12, 75 p. (In Chinese)Google Scholar
Churkin, M., and Carter, C. 1970. Early Silurian graptolites from southeastern Alaska and their correlation with graptolitic sequences in North America and the Arctic. U.S. Geological Survey Professional Paper, 653, 51 p.Google Scholar
Churkin, M., Carter, C., and Eberlin, G. D. 1971. Graptolite succession across the Ordovician-Silurian boundary in south-eastern Alaska. Quarterly Journal of the Geological Society of London, 126:319330.CrossRefGoogle Scholar
Crowther, P. 1981. The fine structure of graptolite periderm. Special Papers in Palaeontology, 26, 119 p.Google Scholar
Crowther, P., and Rickards, R. B. 1977. Cortical bandages and the graptolite zooid. Geologica et Palaeontologica, 11:946.Google Scholar
Davies, K. A. 1929. Notes on the graptolite fauna of the Upper Ordovician and Lower Silurian. Geological Magazine, 66:127.CrossRefGoogle Scholar
Elles, G. L., and Wood, E. M. R. 1901-1918. A monograph of British graptolites. Monograph of the Palaeontographical Society, 539 p.CrossRefGoogle Scholar
Fortey, R. A., and Cooper, R. A. 1986. A phylogenetic classification of the graptoloids. Palaeontology, 29:631654.Google Scholar
Howe, M. 1983. Measurement of thecal spacing in graptolites. Geological Magazine, 120:635638.CrossRefGoogle Scholar
Hundt, R. 1942. Der Schwebeapparat der diprionitischen Graptolithen. Beiträge zur Geologie von Thüringens, 7:7174.Google Scholar
Hutt, J. E. 1974. The Llandovery graptolites of the English Lake District. Part 1. Monograph of the Palaeontographical Society, London, 56 p.Google Scholar
Jones, W. D. V., and Rickards, R. B. 1967. Diplograptus penna Hopkinson 1869, and its bearing on vesicular structures. Paläontologische Zeitschrift, 41:173185.CrossRefGoogle Scholar
Koren', T. N., and Bjerreskov, M. 1997. Early Llandovery monograptids from Bornholm and the southern Urals: taxonomy and evolution. Bulletin of the Geological Society of Denmark, 44:143.CrossRefGoogle Scholar
Koren', T. N., and Rickards, R. B. 1996. Taxonomy and evolution of Llandovery biserial graptoloids from the Southern Urals, Western Kazakhstan. Special Papers in Palaeontology, 54, 49 p.Google Scholar
Koren', T. N., and Sobolevskaya, R. F. 1977. New key sequence of graptolite assemblages across the Ordovician-Silurian boundary (northeast USSR). Doklady Akademii Nauk SSSR, 236:950953. (In Russian)Google Scholar
Koren', T. N., Mikhaylova, N. F., and Tzai, D. T. 1980. Class Graptolithina (Graptolites), p.121170. In Apollonov, M. K., Nikitin, I. F., and Bandaletov, S. M. (eds.), The Ordovician-Silurian Boundary in Kazakhstan. “Nauka” Kazakh SSR. Alma-Ata. (In Russian)Google Scholar
Koren', T. N., Oradovskaya, M. M., Sobolevskaya, R. F., and Chugaeva, M. N. 1979. Atlas of the paleontological plates. Supplement to a Guidebook of the Field Excursion to the Omulev Mountains (Tour VIII). Pacific Science Association. XIV Pacific Science Congress. Magadanskoje Knizhnoje Isdatel'stvo, p. 15. (In Russian)Google Scholar
Koren', T. N., Oradovskaya, M. M., Sobolevskaya, R. F., Pylma, L. J., and Sobolevskaya, R. F. 1983. The Ordovician and Silurian boundary in the northeast of the USSR. Trudy Mezhvedomstvennogo Stratigraficheskogo Komiteta SSSR, 11. Nauka, Leningrad, 205 p. (In Russian)Google Scholar
Lapworth, C. 1873. On an improved classsification of the Rhapdophora. Geological Magazine, 10:500504, 555–560.CrossRefGoogle Scholar
Lapworth, C. 1875. Section Graptoloidea, p.631672. In Hopkinson, J., and Lapworth, C., Descriptions of the graptolites of the Arenig and Llandeilo rocks of St. David's. Quarterly Journal of the Geological Society of London, 31.Google Scholar
Lapworth, C. 1877. On the graptolites of County Down. Proceedings of the Belfast Naturalists’ Field Club, 1:107147.Google Scholar
Lapworth, C. 1880. On new British graptolites. Annals and Magazine of Natural History, 5:149177.CrossRefGoogle Scholar
Legrand, P. 1976. Contribution a l'étude des graptolites du Llandoverien inferieur de l'oued in Djerane (Tassili N'Ajjer oriental, Sahara algerien). Bulletin Societé d'Histoire Naturelle de l’ Afrique du Nord, Alger, 67:141196.Google Scholar
Legrand, P. 1987. Modo de desarrollo del suborden Diplograptina (Graptolithina) en el Ordovícico superior y en el Silúrico. Revista Española de Paleontologia, 2:5964.Google Scholar
Legrand, P. 1998. Approche stratigraphique de l'Ordovicien terminal et du Silurian inferieur du Sahara algerien par l'étude des Diplograptides (graptolites). Thèse de Doctorat d'Etat en Sciences, Université Bordeaux.Google Scholar
Lenz, A. C., and McCracken, A. D. 1982. The Ordovician-Silurian boundary, northern Canadian Cordillera: graptolite and conodont correlation. Canadian Journal of Earth Sciences, 19:13081322.CrossRefGoogle Scholar
Li, Jijin. 1984. Graptolites across the Ordovician-Silurian boundary from Jingxian, Anhui, p.309388. In Nanjing Institute of Geology and Palaeontology, Academia Sinica (eds.), Stratigraphy and Palaeontology of Systemic Boundaries in China, Ordovician-Silurian Boundary (1). Anhui Science and Technology Publishing House.Google Scholar
Li, Jijin, and Meiyu, Ge. 1981. Development and systematic position of akidograptids. Acta Palaeontologica Sinica, 20:225234.Google Scholar
Lin, Yaokun, and Xu, Chen. 1984. Glyptograptus persculptus Zone— the earliest Silurian zone from Yangzi Gorges, China, p. 203232. In Nanjing Institute of Geology and Palaeontology, Academica Sinica (eds.), Stratigraphy and Palaeontology of Systemic Boundaries in China, Ordovician-Silurian Boundary (1). Anhui Science and Technology Publishing House.Google Scholar
Manck, E. 1923. Untersilurische Graptolitenarten der Zone 10 des Obersilurs, ferner Diversograptus n. gen. Natur Jahrgang, 14:282289.Google Scholar
Melchin, M. J. 1989. Llandovery graptolite biostratigraphy and paleobiogeography, Cape Phillips Formation, Canadian Arctic Islands. Canadian Journal of Earth Sciences, 26:17261746.CrossRefGoogle Scholar
Melchin, M. J. 1998. Morphology and phylogeny of some early Silurian ‘diplograptid’ genera from Cornwallis Island, Arctic Canada. Palaeontology, 41:263315.Google Scholar
Melchin, M. J., and Mitchell, C. E. 1991. Late Ordovician extinction in the Graptoloidea, p.143156. In Barnes, C. R., and Williams, S. H. (eds.), Advances in Ordovician Geology. Geological Survey of Canada, Paper 90–9.Google Scholar
Melchin, M. J., McCracken, A. D., and Oliff, F. J. 1991. The Ordovician-Silurian boundary on Cornwallis and Truro islands, Arctic Canada: preliminary data. Canadian Journal of Earth Sciences, 28:18541862.CrossRefGoogle Scholar
Melchin, M. J., Koren', T. N., and Williams, S. H. 1998. Global correlation of the lower part of the earliest Silurian Akidograptus ascensus-Parakidograptus acuminatus Zone, p. 107108. In Gutiérrez-Marco, J. C. and Rábano, I., (eds.), Proceedings of the Sixth International Graptolite Conference of the GWG (IPA) and the 1998 Field Meeting of the International Subcommission on Silurian Stratigraphy (ICS-IUGS), Instituto Tecnológico Geominero de España, Temas Geológico-Mineros, 23.Google Scholar
Mikhaylova, N. F. 1973. Upper Ordovician and Lower Silurian graptolites in Kazakhstan. Trudy Instituta Geologii i Geofiziki Sibirskogo Otdelenia AN SSSR, 47:1419.Google Scholar
Mu, Enzhi. 1988. The Ordovician-Silurian boundary in China, p.117131. In Cocks, L. R. M. and Rickards, R. B. (eds.), A Global Analysis of the Ordovician-Silurian Boundary. Bulletin of the British Museum (Natural History), Geology Series, 43.Google Scholar
Mu, Enzhi, Jijin, Li, Meiyu, Ge, Xu, Chen, Yaokun, Lin, and Yunan, Ni. 1993. Upper Ordovician graptolites of central China region. Palaeontologica Sinica 182, New Series B, 29, 393 p.Google Scholar
Yunan, Ni. 1978. Lower Silurian graptolites from Yichang, western Hubei. Acta Palaeontologica Sinica, 17:387416. (In Chinese)Google Scholar
Nicholson, H. A. 1867. On some fossils of the lower Silurian rocks of the south of Scotland. Geological Magazine, 4:107113.CrossRefGoogle Scholar
Nicholson, H. A. 1868. On the graptolites of the Coniston Flags; with notes on the British species of the genus Graptolites . Quarterly Journal of the Geological Society of London, 24:521545.CrossRefGoogle Scholar
Nicholson, H. A. 1869. On some new species of graptolites. Annals and Magazine of Natural History, 4:231242.CrossRefGoogle Scholar
Obut, A. M. 1965. Silurian graptolites from the Omulevski (Kolimy Basin). Akademiya Nauk SSSR, Siberskoe Otdelenie Institut Geologii i Geofiziki, p. 3346. (In Russian)Google Scholar
Obut, A. M., Sobolevskaya, R. F., and Nikolaev, A. A. 1967. Graptolites and stratigraphy of the lower Silurian along the margins of the Kolyma massif. Akademia Nauk SSSR, Sibirskoje otdelenije, Institut Geologii I Geofiziki. Ministerstvo Geologii SSSR, Nauchno-Issledovatel'sky Institut Geologii Arktiki, 162 p. (In Russian)Google Scholar
Packham, G. H. 1962. Some diplograptids from the British Lower Silurian. Palaeontology, 5:498526.Google Scholar
Pederson, T. B. 1922. Rastritesskiferen på Bornholm. Meddeleser fra Dansk Geologisk Forening 6(11), 29 p.Google Scholar
Perner, J. 1895. Études sur les Graptolites de Bohěme, IIième Partie. Monographie des Graptolites de l'Étage E. Section a. Gerhard, Prague, 25 p.Google Scholar
PŘíbyl, A. 1947. Classification of the genus Climacograptus Hall, 1865. Bulletin International de l'Académie Tchèque des Sciences, Prague, 48:112.Google Scholar
PŘíbyl, A. 1949. A revision of Diplograptidae and Glossograptidae of the Ordovician of Bohemia. Académie Tchèque de Science, Bulletin Internationale, 50:151.Google Scholar
Rickards, R. B. 1970. The Llandovery (Silurian) graptolites of the Howgill Fells, northern England. Palaeontological Society Monograph, 108 p.Google Scholar
Riva, J. 1988. Graptolites at and below the Ordovician-Silurian boundary on Anticosti Island, p.221237. In Cocks, L. R. M. and Rickards, R. B. (eds.), A Global Analysis of the Ordovician-Silurian Boundary. Bulletin of the British Museum (Natural History), Geology Series, 43.Google Scholar
Russel, J. C., Melchin, M. J., and Koren', T. N. 2000. Development, taxonomy, and phylogenetic relationships of species of Paraclimacograptus (Graptoloidea) from the Canadian Arctic and the southern Urals of Russia. Journal of Paleontology, 74:8491.2.0.CO;2>CrossRefGoogle Scholar
Sennikov, N. V. 1976. Graptolites and Lower Silurian stratigraphy of the Gorny Altai. Isdatel'stwo Nauka, Moscow, 274 p. (In Russian)Google Scholar
Štorch, P. 1983. Subfamily Akidograptinae (Graptolithina) from the lowermost Silurian of Bohemia. Věstník Ústrědního ústavu Geologického, 58:295299.Google Scholar
Štorch, P. 1985. Orthograptus s. l. and Cystograptus (Graptolithina) from the Bohemian Lower Silurian. Vestník Ustredního ustavu Geologického, 60:87100.Google Scholar
Štorch, P. 1986. Ordovician-Silurian boundary in the Prague Basin (Barrandian area, Bohemia). Sbornik geologických Ved, Geologie, 41:69103.Google Scholar
Štorch, P. 1988. The Ordovician-Silurian boundary in the Prague Basin, Bohemia, p. 95100. In Cocks, L. R. M. and Rickards, R. B. (eds.), A Global Analysis of the Ordovician-Silurian Boundary. Bulletin of the British Museum (Natural History), Geology Series, 43.Google Scholar
Štorch, P. 1994. Graptolite biostratigraphy of the Lower Silurian (Llandovery and Wenlock) of Bohemia. Geological Journal, 29:137165.CrossRefGoogle Scholar
Štorch, P. 1996. The basal Silurian Akidograptus ascensus-Parakidograptus acuminatus Biozone in peri-Gondwanan Europe: graptolite assemblages, stratigraphical ranges and palaeobiography. Věstník Českeho Geologického Ústavu, 71:177188.Google Scholar
Štorch, P., and Serpagli, E. 1993. Lower Silurian graptolites from the southwestern Sardinia. Bollettino della Societa Paleontologica Italiana, 32:357.Google Scholar
Strachan, I. 1969. A redescription of W. Carruther's type graptolites. Bulletin of the British Museum (Natural History), 17:15205.Google Scholar
Waern, B. 1948. The Silurian strata of the Kullatorp Core, p.337474. In Waern, B., Thorslund, P., and Henningsmoen, G. (eds.), Deep boring through the Ordovician and Silurian strata at Kinnekulle, Vestergotland. Bulletin of the Geological Institute of Upsala, 32.Google Scholar
Williams, S. H. 1983. The Ordovician-Silurian boundary graptolite fauna of Dob's Linn, southern Scotland. Palaeontology, 26:605639.Google Scholar
Yu, Jianhua, Yiting, Fang and Daliang, Zhang. 1988. Lungmachi Formation graptolites from Sanlangpu of Xixiang, southern Shaanxi. Acta Palaeontologica Sinica, 327:150163.Google Scholar