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  • G. CONCENÇO (a1), A. ANDRES (a1), F. SCHREIBER (a1), I. S. MOISINHO (a1), M. C. CORADINI (a1) and M. B. MARTINS (a1)...


We aimed to assess the potential of the characterization of association among weed species as a tool to understand weed occurrence for further supporting long-term management programs. After a sequence of summer crops, which included irrigated rice and sorghum, the experimental area was submitted to subsoiling, limestone was applied, and ryegrass was planted in the winter season. Six months later, an ACCase-inhibitor herbicide was used to select only non-grassweed species. Field survey was carried out on 100 quadrats with 0.5-m width that were randomly sampled. Plant species were organized in 2 × 2 contingency tables. The results of the calculated chi-squares were compared to the respective tables, and results were presented as a paired chi-square matrix. The species–area curve was also obtained. The relative occurrence of species was determined by its frequency and presented as a wordcloud. The network analysis was obtained by using the Fruchterman–Reingold layout. The hypothesis of plant association aiming survival in arable fields was validated. The methodology of plant association based on the chi-square test was applicable to arable fields, where weed species (usually competitor plant types) occur in clusters. From a practical point of view, preference should be given to herbicides that are efficient on most species within a given cluster.


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Adegas, F. S., Oliveira, M. F., Vieira, O. V., Prete, C. E. C., Gazziero, D. L. P. and Voll, E. (2010). Phytosociological survey of weeds in sunflower crop. Planta Daninha 28:705716.
Aldrich, R. J. (1984). Weed-Crop Ecology: Principles in Weed Management. North Scituate: Breton.
Barbour, M. G., Burk, J. H. and Pitts, W. D. (1998). Terrestrial Plant Ecology. Menlo Park: Benjamin/Cummings.
Bastiani, M. O., Lamego, F. P., Agostinetto, D., Langaro, A. C. and Silva, D. C. (2016). Relative competitiveness of soybean cultivars with barnyardgrass. Bragantia 75:435445.
Borsboom, D. and Cramer, A. O. J. (2013). Network analysis: An integrative approach to the structure of psychopathology. Annual Review in Clinical Psychology 9:91121.
Callaway, R. M. (2007). Positive Interactions and Interdependence in Plant Communities. Berlin: Springer.
Chapman, S. B. (1976). Methods in Plant Ecology. Oxford: Blackwell.
Dekker, J. (2009). The Evolutionary Ecology of Weeds and Invasive Plants. Ames: Iowa State University.
Franco, J. J., Agostinetto, D., Langaro, A. C., Perboni, L. T. and Vargas, L. (2017). Relative competitiveness of goosegrass biotypes and soybean crops. Caatinga 30:271277.
Fruchterman, T. M. J. and Reingold, E. M. (1991). Graph drawing by force-directed placement. Software Practice & Experience 21:11291164.
Gurevitch, J., Scheiner, S. M. and Fox, G. A. (2009). Ecologia Vegetal. Porto Alegre: Artmed.
Herben, T., Tackenberg, O. and Klimešová, J. (2016). Reproduction by seed and clonality in plants: Correlated syndromes or independent strategies? Journal of Ecology 104:16961706.
Imperatriz-Fonseca, V. L., Saraiva, A. M. and Jong, D. (2006). Bees as Pollinators in Brazil. Ribeirão Preto: Holos.
Jakelaitis, A., Ferreira, L. R., Silva, A. A., Agnes, E. L., Miranda, G. V. and Machado, A. F. L. (2003). Weed population dynamics under different corn and bean production systems. Planta Daninha 21:7179.
Monquero, P. A. (2014). Aspectos da Biologia e Manejo das Plantas Daninhas. São Carlos: RIMA.
Nicholson, M. (2016). Robert Harding Whittaker and the individualistic hypothesis. Bulletin of the Ecological Society of America 97:154162.
Paterno, G. B., Siqueira, F. J. A. and Ganade, G. (2016). Species-specific facilitation, ontogenetic shifts and consequences for plant community succession. Journal of Vegetation Science 27:606615.
Santos, R. N. V., Rodrigues, A. A. C., Silva, M. R. M., Correa, M. J. P. and Mesquita, M. L. R. (2017). Phytosociology and weed interference in okra under organic cropping system. African Journal of Agricultural Research 12:251259.
Schöb, C., Macek, P., Pistón, N., Kilkvidze, Z. and Pugnaire, F. I. (2017). A trait-based approach to understand the consequences of specific plant interactions for community structure. Journal of Vegetation Science 28:696704.
Singh, H. P., Batish, D. R. and Hohli, R. K. (2006). Handbook of Sustainable Weed Management. New York: CRC Press.
Spotswood, E. N., Mariotte, P., Farrer, E. C., Nichols, L. and Suding, K. N. (2017). Separating sources of density-dependent and density-independent establishment limitation in invading species. Journal of Ecology 105:436444.
Tuffi-Santos, L. D., Santos, I. C., Oliveira, C. H., Santos, M. V., Ferreira, F. A. and Queiroz, D. S. (2004). Phyto-sociological assessment of degraded pastures under flooded low land conditions. Planta Daninha 22:343349.
Voinov, V., Nikulin, M. and Balakrishnan, N. (2013). Chi-Squared Goodness of Fit Tests with Applications. Amsterdam: Elsevier.
Whittaker, R. H. (1960). Vegetation of Siskiyou mountains, Oregon and California. Ecological Monographs 30:279338.
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  • G. CONCENÇO (a1), A. ANDRES (a1), F. SCHREIBER (a1), I. S. MOISINHO (a1), M. C. CORADINI (a1) and M. B. MARTINS (a1)...


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