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Late Ordovician trilobites from the Mayatas Formation, Atansor area, north-central Kazakhstan

Published online by Cambridge University Press:  07 September 2017

L. M. E. McCobb
Affiliation:
Department of Natural Sciences, National Museum Wales, Cathays Park, Cardiff CF10 9NP, UK. E-mail: lucy.mccobb@museumwales.ac.uk; leonid.popov@museumwales.ac.uk
L. E. Popov
Affiliation:
Department of Natural Sciences, National Museum Wales, Cathays Park, Cardiff CF10 9NP, UK. E-mail: lucy.mccobb@museumwales.ac.uk; leonid.popov@museumwales.ac.uk

Abstract

Two trilobite faunas of Late Ordovician (Katian) age are described from the Mayatas Formation in the Stepnyak region of north-central Kazakhstan. The older, oligotaxic fauna derives from flanks of a carbonate build-up, and is dominated by numerous Sphaerexochus specimens. Amphilichas is also relatively common, with Pliomerina and indeterminate asaphids present as rare components. The overlying unit of siliceous argillites contains a different assemblage, representing the raphiophorid biofacies and comprising seven genera. The poorly preserved fauna is dominated by blind trilobites (a new genus of trinucleid, the three-segmented raphiophorid Pseudampyxina, Malongullia?, Lonchodomas and Arthrorhachis) and at least two species of large-eyed Telephina, suggesting that they occupied the disphotic zone in deep water offshore. A single cranidium of the odontopleurid Primaspis is also present. The trinucleid, Iputaspis stepnyakensis gen. et sp. nov., has an unusual pit arrangement, with E1 and E2 aligned in sulci and all I arcs irregularly arranged. The Atansor area is located within the Stepnyak tectonostratigraphical unit, which probably represented an Ordovician active margin of the Kalmykkol–Kokchetav Microplate. Some of the genera represented in the faunas have affinities with Australia and South China and, also, there is a possible link to European peri-Gondwana.

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Copyright © The Royal Society of Edinburgh 2017 

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References

5. References

Ahlberg, P. 1995. Telephinid trilobites from the Ordovician of Sweden. Palaeontology 38(2), 259–85.Google Scholar
Angelin, N. P. 1854. Palaeontologica Scandinavica. Pars 1. Crustacea Formayionis Transitionis. Fasc. 1. Palaeontologica Suecica, 124.Google Scholar
Apollonov, M. K. 1974. [Ashgill trilobites of Kazakhstan.] Alma-Ata: Nauka. 134 pp. [In Russian.]Google Scholar
Balashova, E. A. 1960. [Trilobites of the Middle and Upper Ordovician and the Lower Silurian of Eastern Taimyr.] Leningrad University. 95 pp., 6 pls. [In Russian.]Google Scholar
Balashova, E. A. 1961. [Discovery of a new trilobite in the Glauconite Series of the Baltic region.] Paleontological Journal 1961(3), 129–32. [In Russian.]Google Scholar
Barrande, J. 1846. Notice préliminaire sur le système Silurien et les Trilobites de Bohême. Leipzig: Hirschfeld. 97 pp.Google Scholar
Barrande, J. 1852. Systême Silurien du centre de la Bohême, 1ère Partie. Récherches paléontologiques. Vo. 1, Crustacés: Trilobites. Prague and Paris.Google Scholar
Bazhenov, M. L., Levashova, N. M., Degtyarev, K. E, Van der Voo, R., Abrajevitch, A. V. & McCausland, P. J. A. 2012. Unraveling the early–middle Paleozoic paleogeography of Kazakhstan on the basis of Ordovician and Devonian paleomagnetic results. Gondwana Research 22, 974–91.Google Scholar
Beyrich, E. 1845. Ueber einige böhmische Trilobiten. Berlin: Reimer. 47 pp.Google Scholar
Bruton, D. L. 1966. A revision of the Swedish Ordovician Odontopleuridae (Trilobita). Bulletin of the Geological Institutions of the University of Uppsala 43, 140.Google Scholar
Bruton, D. L. 1968. A revision of the Odontopleuridae (Trilobita) from the Palaeozoic of Bohemia. Skrift Norske Vidensk-Akad Oslo. I. Mat.-Naturv. Klasse: NY Serie 24, 173.Google Scholar
Bruton, D. L. & Høyberget, M. 2006. A reconstruction of Telephina bicuspis, a pelagic trilobite from the Middle Ordovician of the Oslo region, Norway. Lethaia 39, 359–64.Google Scholar
Budil, P., Fatka, O., Kolář, P. & David, M. 2011. Arthrorhachis Hawle & Corda, 1984 (Agnostida) in the Prague Basin (Barrandian area, Czech Republic) revisited. Bulletin of Geosciences 86(4), 707–24.Google Scholar
Burmeister, H. 1843. Die Organisation der Trilobiten aus ihren lebenden Verwandten entwickelt; nebst einer systematischen Uebersicht aller zeither beshreibenen Arten, ixii. Berlin: Ray Society. 147 pp.Google Scholar
Burrett, C., Stait, B. & Laurie, J. 1983. Trilobites and microfossils from the Middle Ordovician of Surprise Bay, southern Tasmania, Australia. Memoirs of the Association of Australasian Palaeontologists 1, 177–93.Google Scholar
Carlucci, J. R., Westrop, S. R. & Amati, L. 2010. Tetralichine trilobites from the Upper Ordovician of Oklahoma and Virginia and phylogenetic systematics of the Tetralichini. Journal of Paleontology 84, 1099–120.Google Scholar
Chu, C., Lin, H. & Zhang, Z. 1979. Trilobita. In Nanjing Institute of Geology and Palaeontology, Qinghai Institute of Geology (eds) Palaeontological Atlas of Northwest China, Qinghai Province, Volume 2, 81116. Beijing: Geological Publishing House.Google Scholar
Chugaeva, M. N. 1958. [Trilobites from the Ordovician of Chu-Ili Range.] Trudy geologicheskogo Instituta Akademii Nauk SSSR 9, 5138. [In Russian.]Google Scholar
Conway, T. M. & Botting, J. P. 2012. Description and ecology of a new Middle Ordovician (Llanvirn) odontopleurid trilobite from the Builth Inlier of mid-Wales, with a review of the genus Meadowtownella. Geological Magazine 149(3), 397411.Google Scholar
Dean, W. T. 1973. The lower Palaeozoic stratigraphy and faunas of the Taurus Mountains near Beysehir, Turkey III. The trilobites of the Sobova Formation (Lower Ordovician). Bulletin of the British Museum (Natural History) 24, 281348.Google Scholar
Dean, W. T. & Zhou, Z. 1988. Upper Ordovician trilobites from the Zap Valley, southeast Turkey. Palaeontology 31, 621–49.Google Scholar
Degtyarev, K. E. 2012. [Tectonic evolution of the Early Palaeozoic island arc systems and continental core formation of Kazakhstanian Caledonides.] Transactions of the Geological Institute, Russian Academy of Sciences 602. Moscow: Geos Publishing House. 289 pp. [In Russian.]Google Scholar
Dobretsov, N. K., Buslov, M. M., Zhimulev, F. I., Travin, A. V. & Zayachkovskii, A. A. 2006. [Vendian–Early Ordovician geodynamic evolution and model for exhumation of ultrahigh- and high-pressure rocks from the Kokchetav subduction-collision zone.] Geologiya I geofizika 47, 428–44. [In Russian.]Google Scholar
Edgecombe, G. D., Banks, M. R. & Banks, D. M. 1999a. Upper Ordovician Phacopida (Trilobita) from Tasmania. Alcheringa 23, 235–57.Google Scholar
Edgecombe, G. D., Chatterton, B. D. E., Waisfeld, B. G. & Vaccari, N. E. 1999b. Ordovician pliomerid and prosopiscid trilobites from Argentina. Journal of Paleontology 73, 1144–54.Google Scholar
Edgecombe, G. D., Banks, M. R. & Banks, D. M. 2004. Late Ordovician trilobites from Tasmania: Styginidae, Asaphidae and Lichidae. Memoirs of the Association of Australasian Palaeontologists 30, 5977.Google Scholar
Edgecombe, G. D. & Webby, B. D. 2007. Ordovician trilobites with eastern Gondwanan affinities from central-west New South Wales and Tasmania. Memoirs of the Association of Australasian Palaeontologists 34, 255–81.Google Scholar
Fatka, O., Lerosey-Aubril, R., Budil, P. & Rak, Š. 2013. Fossilised guts in trilobites from the Upper Ordovician Letná Formation (Prague Basin, Czech Republic). Bulletin of Geosciences 88(1), 95104. DOI 10.3140/bull.geosci.1329Google Scholar
Foerste, A. F. 1919. Notes on Isotelus, Acrolichas, Calymene and Encrinurus. Bulletin of the scientific labs of Denison University 19, 681.Google Scholar
Fortey, R. A. 1975a. Early Ordovician trilobite communities. Fossils and Strata 4, 331–52.Google Scholar
Fortey, R. A. 1975b. The Ordovician trilobites of Spitsbergen. II. Asaphidae, Nileidae, Raphiophoridae and Telephinidae of the Valhallfonna Formation. Norsk Polarinstitutt Skrifter 162, 1207.Google Scholar
Fortey, R. A. 1980. The Ordovician trilobites of Spitsbergen. III. Remaining trilobites of the Vallhallfonna Formation. Norsk Polarinstitutt Skrifter 171, 1163.Google Scholar
Fortey, R. A. 1997. Late Ordovician trilobites from southern Thailand. Palaeontology 40, 397449.Google Scholar
Fortey, R. A. & Owens, R. M. 1975. Proetida: a new order of trilobites. Fossils and Strata 4, 227–39.Google Scholar
Ghobadi Pour, M., Popov, L. E. & Vinogradova, E. V. 2009. Middle Ordovician (late Darriwilian) trilobites from the northern Betpak-Dala Desert, central Kazakhstan. Memoirs of the Association of Australasian Palaeontologists 37, 327–49.Google Scholar
Ghobadi Pour, M., Popov, L. E., McCobb, L. M. E. & Percival, I. G. 2011. New data on the Late Ordovician trilobite faunas of Kazakhstan: implications for biogeography of tropical peri-Gondwana. In: Gutiérrez-Marco, J. C., Rábano, I. & García-Bellido, D. (eds), Ordovician of the World. Cuadernos del Museo Geominero, 14, 171–77. Madrid: Instituto Geológico y Minero de España.Google Scholar
Han, N. 2001. The eyes of Ordovician trilobite Telephina convexa Lu. Acta Palaeontologica Sinica 40(3), 399408.Google Scholar
Harrington, H. J. & Leanza, A. F. 1957. Ordovician trilobites of Argentina. The University of Kansas Paleontological Contributions: Special Publication 1. Lawrence, Kansas: University of Kansas Press. 276 pp.Google Scholar
Hawle, I. & Corda, A. J. S. 1847. Prodrom einer Monographie der böhmischen Trilobiten. Abhandlungen Koeniglichen Boehmischen Gesellschaft der Wissenschaften. Prague: J. G. Calve. 176 pp.Google Scholar
Jaekel, O. 1909. Über die Agnostiden. Zeitschrift der Deutschen Geologischen gesellschaft 61, 380401.Google Scholar
Jell, P. A. & Adrain, J. M. 2002. Available generic names for trilobites. Memoirs of the Queensland Museum 48(2), 331553.Google Scholar
Kobayashi, T. 1934. The Cambro-Ordovician Formations and Faunas of South Chosen. Palaeontology. Pt. I. Middle Ordovician faunas. Journal of the Faculty of Science Imperial University of Tokyo, Section 2 3(8), 329519.Google Scholar
Kobayashi, T. & Hamada, T. 1971. Silurian trilobites from the Langkawi Islands, West Malaysia, with notes on the Dalmanitidae and Raphiophoridae. Geology and Palaeontology of Southeast Asia 9, 87134.Google Scholar
Kolobova, I. M. 1985. [On a new raphiophorid genus (Trilobita) from the Middle Ordovician of Kazakhstan.] Ezhegodnik Vsesoyuznogo Paleontologicheskogo Obshchestva 28, 168–75. [In Russian.]Google Scholar
Koroleva, M. N. 1954. [Limestones of the Lake Atansor area in northeastern Kazakhstan.] Geologiya, Gornoye Delo i Metallurgiya 9, 133–36. [In Russian.]Google Scholar
Koroleva, M. N. 1959. [New genera of trilobites from the Middle and Upper Ordovician of northern Kazakhstan.] Doklady Akademii Nauk SSSR 124, 1313–16. [In Russian.]Google Scholar
Koroleva, M. N. 1965. [New Ordovician trilobites from Central Kazakhstan.] Ezhegodnik Vsesoyuznogo Paleontologicheskogo Obshchestva 17, 148–73. [In Russian.]Google Scholar
Koroleva, M. N. 1982. [Ordovician trilobites of north-eastern Kazakhstan.] Moscow: Nedra. 192 pp. [In Russian.]Google Scholar
Koroleva, M. N. 1997. [Trilobites of genus Sphaerexochus from Kazakhstan.]. Almaty: TOO International Geophysics. 18 pp. [In Russian.]Google Scholar
Kröner, A., Hegner, E., Lehmann, B., Heinhorst, J., Wingate, M. T. D., Liu, D. & Ermelov, P. 2008. Palaeozoic arc magmatism in the Central Asian Orogenic Belt of Kazakhstan: SHRIMP zircon ages and whole-rock Nd isotopic systematics. Journal of Asian Earth Sciences 32, 118–30.Google Scholar
Lu, Y. 1975. Ordovician trilobite faunas of central and south-western China. Palaeontologica Sinica 11, 319–39.Google Scholar
Marek, L. 1952. Contribution to the stratigraphy and fauna of the uppermost part of the Králův Dvůr Shales (Ashgillian). Sborník Úst"edního Ústavu Geologckého 19, 429–55.Google Scholar
M'Coy, F. 1846. A synopsis of the Silurian fossils of Ireland. Dublin: At the University Press by M. H. Gill. 72 pp.Google Scholar
M'Coy, F. 1849. On the classification of some British fossil Crustacea with notices of some forms in the University collection at Cambridge. Annals and Magazine of Natural History 4(2), 161–79, 330–35, 392–414.Google Scholar
Meek, F. B. & Worthen, A. H. 1868. Palaeontology. Geological Survey of Illinois 3, 289572.Google Scholar
Melnikova, L. M. 1980. [Some Early Ordovician Ostracodes of the southern Urals.] Paleontological Journal 1980(3), 7176. [In Russian.]Google Scholar
Melnikova, L. M. 1986. Ordovikskiye ostrakody Kazakhstana. Trudy Paleontologicheskogo Instituta 218, 1103.Google Scholar
Mergl, M. 2014. Odontopleurid trilobites of the Katian/Hirnantian boundary interval in the Prague Basin (Bohemia). Bulletin of Geosciences 89(2), 401–12.Google Scholar
Moore, R. C. 1959. Treatise on Invertebrate Paleontology, Part O, Arthropoda 1. Boulder, Colorado & Lawrence, Kansas: Geological Society of America and University of Kansas Press. 560 pp.Google Scholar
Nazarov, B. B. & Popov, L. E. 1980. [Stratigraphy and fauna of Ordovician siliceous-carbonate deposits of Kazakhstan.] Trudy Geolgicheskogo Instituta Akademii Nauk SSSR 331, 1190. [In Russian.]Google Scholar
Nielsen, A. T. 1997. A review of Ordovician agnostid genera (Trilobita). Transactions of the Royal Society of Edinburgh: Earth Sciences 87(for 1996), 463501.Google Scholar
Nikitin, I. F. 1972. [Ordovician of Kazakhstan. Part 1. Stratigraphy.] Alma-Ata: Nauka. 242 pp. [In Russian.]Google Scholar
Nikitin, I. F. (ed.) 1991. [Decisions of the Third Stratigraphical Conference on the Precambrian and Phanerozoic of Kazakhstan, Almaty, 1986. Part I. Precambrian and Palaeozoic.] St. Petersburg–Alma Ata: VSEGEI. 148 pp. [In Russian].Google Scholar
Öpik, A. A. 1937. Trilobiten aus Estland. Acta et Commentationes Universitatis Tartuensis (Dorpatensis) A 32(3), 1163.Google Scholar
Owen, A. W. & Parkes, M. A. 2000. Trilobite faunas of the Duncannon Group: Caradoc stratigraphy, environments and palaeobiogeography of the Leinster Terrane, Ireland. Palaeontology 43(2), 219–69.Google Scholar
Phleger, F. B. 1936. Lichadian trilobites. Journal of Paleontology 10, 593615.Google Scholar
Popov, L. E. 2000. Late Ordovician (Caradoc) linguliformean microbrachiopods from north-central Kazakhstsan. Alcheringa 24, 257–75.Google Scholar
Popov, L. E., Bassett, M. G., Zhemchuzhnikov, V. G., Holmer, L. E. & Klishevich, I. A. 2009. Gondwanan faunal signatures from Early Palaeozoic terranes of Kazakhstan and Central Asia: evidence and tectonic implications. Geological Society, London, Special Publication 325, 2364.Google Scholar
Prantl, F. & Pribyl, A. 1949. A study of the superfamily Odontopleuracea nov. superfam. (Trilobita). Rozpravy Státniho Geologického Ústavu ČSR 12, 1221.Google Scholar
Pribyl, A., Vaněk, J. & Pek, I. 1985. Phylogeny and taxonomy of Family Cheiruridae (Trilobita). Acta Universitatis Palackianae Olomucensis Facultas rerum naturalium. Geographica-Geologica 83, 107–93.Google Scholar
Pribyl, A. & Vaňek, J. 1965. Neue Trilobiten des böhmischen Ordoviziums. Vestník Ústředního Ústavu Geologického 40, 277–82.Google Scholar
Qui, H., Lu, Y., Zhu, Z., Bi, D., Lin, T., Zhou, Z., Zhang, Q., Qian, Y., Ju, T., Han, N. & Wei, X. 1983. Palaeontological Atlas of East China. Part 1: Early Palaeozoic. Nanjing Institute of Geology and Mineral Resources. Beijing: Geological Publishing House. 637 pp; 176 pls.Google Scholar
Ramsköld, L. & Chatterton, B. D. E. 1991. Revision and subdivision of the polyphyletic ‘Leonaspis’ (Trilobita). Transactions of the Royal Society of Edinburgh: Earth Sciences 82, 333–71.Google Scholar
Raymond, P. E. 1905. Trilobites of the Chazy Limestone. Annals of the Carnegie Museum 3, 328–86.Google Scholar
Raymond, P. E. 1913. Notes on some new and old trilobites in the Victoria Memorial Museum, Canadian Geological Survey (Ottawa). Bulletin of the Victoria Memorial Museum 1, 3339.Google Scholar
Raymond, P. E. 1920. Some new Ordovician trilobites. Bulletin of the Museum of Comparative Anatomy, Harvard 64, 272–96.Google Scholar
Reed, F. R. C. 1917. Ordovician and Silurian fossils from Yun-nan. Memoirs of the Geological Suvery of India. Palaeontologia Indica, N. S. 6, 184.Google Scholar
Richter, R. & Richter, E. 1917. Über die Einteilung der Familie Acidaspidae und über einige ihrer devonishcen Vertreter. Zentralblatt für Mineralogie, Geologie und Paläontologie 1917, 462–72.Google Scholar
Salter, J. W. 1864. The British trilobites from the Cambrian, Silurian and Devonian formations. Part I. (Devonian and Silurian). Monographs of the Palaeontographical Society 67(16), 180. London: The Palaeontographical Society.Google Scholar
Sars, M. 1835. Über einige neue oder unvollständig bekannte Trilobiten. Isis 28, 333–43.Google Scholar
Şengör, A. M. C. & Natal'in, B. A. 1996. Paleotectonics of Asia: fragments of a synthesis. In Yin, A. & Harrison, M. (eds) The Tectonic Evolution of Asia, 486640. Cambridge, New York, Port Chester, Melbourne, Sydney: Cambridge University Press. xii + 666 pp.Google Scholar
Siveter, D. J. 1989. Silurian trilobites from the Annascaul inlier, Dingle Peninsula, Ireland. Palaeontology 32, 109–61.Google Scholar
Šnajdr, M. 1984. Bohemian Ordovician Odontopleuridae (Trilobita). Sbornik Geologických Ved Palaeontology 26, 4782.Google Scholar
Sun, Z. 1984. [Trilobita.] In [The Palaeontological Atlas of Hubei Province], 328422. Wuhan: Hubei Science and Technology Press. [In Chinese.]Google Scholar
Thomas, A. T. & Holloway, D. J. 1988. Classification and phylogeny of the trilobite order Lichida. Philosophical Transactions of the Royal Society, London B321, 179262.Google Scholar
Torsvik, T. H. & Cocks, L. R. M. 2009. The Lower Palaeozoic palaeogeographical evolution of the northeastern and eastern peri-Gondwanan margin from Turkey to New Zealand. Geological Society, London, Special Publications 325, 321.Google Scholar
Tripp, R. P. 1957. The classification and evolution of the superfamily Lichacea (Trilobita). Geological Magazine 94, 104–22.Google Scholar
Tripp, R. P., Zhou, Z. & Pan, Z. 1989. Trilobites from the Upper Ordovician Tangton Formation, Jiangsu Province, China. Transactions of the Royal Society of Edinburgh: Earth Sciences, 80, 2568.Google Scholar
Vaccari, N. E., Waisfeld, B. G., Chatterton, B. D. E. & Edgecombe, G. D. 2006. New raphiophorid trilobites from the Ordovician of Argentina and their biogeographic implications. Memoirs of the Association of Australasian Palaeontologists 32, 353–74.Google Scholar
Warburg, E. 1925. The trilobites of the Leptaena Limestone in Dalarna. Bulletin of the Geological Institution, University of Uppsala 17, 1446.Google Scholar
Webby, B. D. 1971. The trilobite Pliomerina Chugaeva from the Ordovician of New South Wales. Palaeontology 14, 612–22.Google Scholar
Webby, B. D. 1974. Upper Ordovician trilobites from central New South Wales. Palaeontology 17, 203–52.Google Scholar
Webby, B. D., Moors, H. T. & McLean, R. A. 1970. Malongullia and Encrinuraspis, New Ordovician trilobites from New South Wales, Australia. Journal of Paleontology 44, 881–87.Google Scholar
Weber, V. N. 1948. [Silurian Trilobites of the USSR. 1: Lower Silurian trilobites.] Monograph on the Palaeontology of the USSR 69(1), 1113. [In Russian.]Google Scholar
Whittington, H. B. 1956. Silicified Middle Ordovician trilobites: the Odontopleuridae. Bulletin of the Museum of Comparative Anatomy, Harvard University 114(5), 155288.Google Scholar
Zhang, W. 1980. On the Miomera and Polymera (Trilobita). Scientia Sinica 10, 9961004.Google Scholar
Zhang, W. & Fan, J. 1960. Ordovician and Silurian trilobites of the Chilian Mountains. In Yin, T. (ed.) Geological Gazetteer of the Chilian Mountains 4, 83148, pls 110. Beijing: Science Press. 160 pp.Google Scholar
Zhang, W. & Jell, P. A. 1987. Cambrian trilobites of North China: Chinese Cambrian trilobites housed in the Smithsonian Institution. Beijing: Science Press. xvi + 459 pp; 125 pl.Google Scholar
Zhou, Z., Li, J. & Qu, X. 1982. Trilobita. In Xian Institute of Geology and Mineral Resources (eds) Palaeontological Atlas of Northwest China, Shaanxi–Gansu–Ningxia Part 1: Precambrian-Lower Palaeozoic, 215–94. Beijing: Geological Publishing House.Google Scholar
Zhou, Z., Webby, B. D. & Yuan, W. 1995. Ordovician trilobites from the Yingan Formation of northwestern Tarim, Xinjiang, northwestern China. Alcheringa 19, 4772.Google Scholar
Zhou, Z. & Fortey, R. A. 1986. Ordovician trilobites from north and northeastern China. Palaeontographica Abteilung A 192, 157210.Google Scholar
Zhou, Z. Q. & Zhou, Z. Y. 2006. Late Ordovician trilobites from the Zhusilenghaierhan area, Ejin Banner, western Inner Mongolia, China. Memoirs of the Association of Australasian Palaeontologists 32, 383411.Google Scholar