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True flies (Insecta: Diptera) from the late Eocene insect limestone (Bembridge Marls) of the Isle of Wight, England, UK

Published online by Cambridge University Press:  04 October 2019

Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Ul. Sławkowska 17, 31-016 Kraków, Poland. Email:
Department of Natural Sciences, National Museums Scotland, Chambers St., Edinburgh EH1 1JF, UK.
Department of Biology, Faculty of Sciences II, Lebanese University, Fanar – Matn, P. O. box 26110217, Lebanon.
Paleontological Institute, Russian Academy of Sciences, Profsoyuznaya Str. 123, 117868 Moscow, Russia.
Department of Invertebrate Zoology, University of Gdańsk, Pilsudskiego 46, 81-378 Gdynia, Poland.
Senckenberg Research Institute and Museum, Research station Grube Messel, Markstraße 35, D-64409 Messel, Germany.
André NEL
CNRS UMR 5202, Muséum National d'Histoire Naturelle, CP 50, Entomologie, 45 rue Buffon, F-75005, Paris, France.
François-Marie COLLOMB
CNRS UMR 5202, Muséum National d'Histoire Naturelle, CP 50, Entomologie, 45 rue Buffon, F-75005, Paris, France.
CNRS UMR 5202, Muséum National d'Histoire Naturelle, CP 50, Entomologie, 45 rue Buffon, F-75005, Paris, France.
School of Earth and Environmental Sciences, University of Portsmouth, Burnaby Road, Portsmouth PO1 3QL, UK.
*Corresponding author


The Diptera fauna from the late Eocene of the Isle of Wight (Bembridge Marls) is studied including redescriptions of formerly described material. The fauna includes the following taxa: Anisopodidae – one species; Bibionidae – 11 species; Ceratopogonidae – one described and two unidentified species; Chironomidae – undetermined species of three subfamilies; Culicidae – four species; Cylindrotomidae – one species; Dixidae – one species; Keroplatidae – one described and four unidentified species; Limoniidae – 31 species; Mycetophilidae – 14 species; Psychodidae – two species; Scatopsidae – two species; Sciaridae – three species; Simuliidae – an unnamed species; Tipulidae – nine species. Diptera Brachycera: Agromyzidae – unidentified species; Anthomyzidae – two species; Asilidae – two species; Bombyliidae – one species; Chloropidae – one species; Dolichopodidae – unidentified species; Empididae – unidentified species; Ephydridae – two species; Heleomyzidae – unidentified species; Lauxaniidae – unidentified species; Otitidae – one species; Phoridae – unidentified species; Rhagionidae – two species; Stratiomyidae – two species; Syrphidae – unidentified species; Tabanidae – one species. Following new taxa are described: Leptotarsus (Longurio) fragmentatus Krzemiński, n. sp., Leptotarsus (Longurio) wegiereki Krzemiński, n. sp., Tipula (s. lato) anglicana Krzemiński, n. sp. (Tipulidae); Cyttaromyiarossi Krzemiński, n. sp. (Cylindrotomidae); Gurnardia corami Krzemiński, n. gen, n. sp., Pilaria hooleyi Krzemiński, n. sp., Pilaria volodii Krzemiński, n. sp., Cheilotrichia (Cheilotrichia) duplicata Krzemiński, n. sp., Cheilotrichia (Empeda) szwedoi Krzemiński, n. sp., Symplecta (Psiloconopa) gurnetensis Krzemiński, n. sp., Orimarga (Orimarga) lenae Krzemiński, n. sp., Dicranomyia (Dicranomyia) azari Krzemiński, n. sp., Dicranomyia (Dicranomyia) fasciata Krzemiński, n. sp., Helius (Helius) edmundi Krzemiński, n. sp., Helius (Helius) popovi Krzemiński, n. sp., Dicranoptycha staryi Krzemiński, n. sp. (Limoniidae); Wightipsychoda Azar, n. gen. (Psychodidae); Dilophus andrewrossi Nel, Colomb & Waller, n. sp., Bibiodes massiliensis Nel, Colomb & Waller, n. sp. (Bibionidae); Azana cockerelli Blagoderov, n. sp., Aglaomyia vectis Blagoderov, n. sp., Dziedzickia oligocenica Blagoderov, n. sp., Palaeoempalia saxea Blagoderov, n. sp., Mycomyahoolei Blagoderov, n. sp., Leia gurnardensis Blagoderov, n. sp. (Mycetophilidae); Regmocleminahaennii Krzemiński, n. sp. (Scatopsidae); Sylvicola problematica Krzemiński, n. sp. (Anisopodidae); Palaeoberidops barkeri Nicholson, n. gen., n. sp. (Stratiomyidae). The Diptera fauna and palaeohabitat of the Insect Bed is discussed.

Copyright © The Royal Society of Edinburgh 2019 

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